The DOI matches its Crossref record; 225 of the 225 references it cites were found in a registry or an index.
How deep is the brain? The shallow brain hypothesis
Open accessAbstract
Deep learning and predictive coding architectures commonly assume that inference in neural networks is hierarchical. However, largely neglected in deep learning and predictive coding architectures is the neurobiological evidence that all hierarchical cortical areas, higher or lower, project to and receive signals directly from subcortical areas. Given these neuroanatomical facts, today's dominance of cortico-centric, hierarchical architectures in deep learning and predictive coding networks is highly questionable; such architectures are likely to be missing essential computational principles the brain uses. In this Perspective, we present the shallow brain hypothesis: hierarchical cortical processing is integrated with a massively parallel process to which subcortical areas substantially contribute. This shallow architecture exploits the computational capacity of cortical microcircuits and thalamo-cortical loops that are not included in typical hierarchical deep learning and predictive coding networks. We argue that the shallow brain architecture provides several critical benefits over deep hierarchical structures and a more complete depiction of how mammalian brains achieve fast and flexible computational capabilities.
Overview
Integrity signals
Registry
Cites
Statistics
Verification
Claims the paper makes
Claims other papers make about this one
Each is checked against this paper’s own text: does it say what the citing paper says it does?
In the citing paper
“In the typical mammalian brain, however, recurrent loops between the subcortex and regions of the cortex are a vital and almost universal feature of processing [85].”
From Phenomenal interface theory: a model for basal consciousness (2025) · cites this paper as [85]
Citations
Cites
Deep Residual Learning for Image Recognition
He, K., Zhang, X., Ren, S., & Sun, J. (2016) · 2016 IEEE Conference on Computer Vision and Pattern Recognition (CVPR) · doi:10.1109/cvpr.2016.90 · Cited by 235,474
LeCun, Y., Bengio, Y., & Hinton, G. (2015) · Nature · doi:10.1038/nature14539 · Cited by 84,126
ImageNet Large Scale Visual Recognition Challenge
Russakovsky, O., Deng, J., Su, H., Krause, J., Satheesh, S., Ma, S., Huang, Z., Karpathy, A., Khosla, A., Bernstein, M., et al. (2015) · International Journal of Computer Vision · doi:10.1007/s11263-015-0816-y · Cited by 39,380
Receptive fields, binocular interaction and functional architecture in the cat’s visual cortex
Hubel, D. H., & Wiesel, T. N. (1962) · The Journal of Physiology · doi:10.1113/jphysiol.1962.sp006837 · Cited by 13,680
An Integrative Theory of Prefrontal Cortex Function
Miller, E. K., & Cohen, J. D. (2001) · Annual Review of Neuroscience · doi:10.1146/annurev.neuro.24.1.167 · Cited by 12,803
Parallel organization of functionally segregated circuits linking basal ganglia and cortex
Alexander, G. E., DeLong, M. R., & Strick, P. L. (1986) · Annual Review of Neuroscience · doi:10.1146/annurev.ne.09.030186.002041 · Cited by 8,527
Show, Attend and Tell: Neural Image Caption Generation with Visual Attention
Xu, K., Ba, J., Kiros, R., Cho, K., Courville, A., Salakhutdinov, R., Zemel, R., & Bengio, Y. (2015) · arXiv · doi:10.48550/arxiv.1502.03044 · Cited by 8,248
The free-energy principle: a unified brain theory?
Friston, K. (2010) · Nature Reviews Neuroscience · doi:10.1038/nrn2787 · Cited by 8,120 · cited 1×
“This idea of hierarchical Bayesian inference has been the dominant framework in neuroscience over the last decade 17,40–43 .”
BackgroundA robust layered control system for a mobile robot
Brooks, R. (1986) · IEEE Journal on Robotics and Automation · doi:10.1109/jra.1986.1087032 · Cited by 7,799 · cited 2×
“Although the cortical hierarchy is useful for representing abstract features in their context, we propose a processing regime, the shallow brain hypothesis, whereby the entire cortex is essentially one giant layer providing contextualized input to subcortical motor and premotor centres in the…”
BackgroundDistributed hierarchical processing in the primate cerebral cortex
Felleman, D. J., & Van Essen, D. C. (1991) · Cerebral Cortex · doi:10.1093/cercor/1.1.1 · Cited by 7,783
Research works printed but not linked: 6
- Minsky, M. & Papert, S. Perceptrons; An Introduction to Computational Geometry (MIT Press, 1969).
- Cohen, N., Sharir, O. & Shashua, A. On the expressive power of deep learning: a tensor analysis. In 29th Annual Conference on Learning Theory (eds. Feldman, V., Rakhlin, A. & Shamir, O.) 698–728 (2016).
- Von Neumann, J. The Computer and the Brain (Yale Univ. Press, 1958).
- Haider, P., Ellenberger, B., Kriener, L., Jordan, J., Senn, W. & Petrovici, M. A. Latent equilibrium: a unified learning theory for arbitrarily fast computation with arbitrarily slow neurons. Adv. Neural Inf. Process. Syst. 34, 17839–17851 (2021).
- Ruis, L., Andreas, J., Baroni, M., Bouchacourt, D. & Lake, B. M. A benchmark for systematic generalization in grounded language understanding. In Proc. 34th Int. Conf. Neural Information Processing Systems (eds. Larochelle, H. et al.) 19861–19872 (Curran, 2020).
- Lake, B. M. & Baroni, M. Generalization without systematicity: on the compositional skills of sequence-to-sequence recurrent networks. In Int. Conf. Machine Learning (eds. Dy, J. & Krause, A.) 2879–2888 (2018).
Cited by
Sorted by citations
An integrative, multiscale view on neural theories of consciousness
Storm, J. F., Klink, P. C., Aru, J., Senn, W., Goebel, R., Pigorini, A., Avanzini, P., Vanduffel, W., Roelfsema, P. R., Massimini, M., et al. (2024) · Neuron · doi:10.1016/j.neuron.2024.02.004 · Cited by 114
“The next aggregate levels are local (within-area) recurrent processing in cortical microcircuits (C, upper left; from Suzuki et al. 138 ) and cortico-thalamic loops (C, HO: higher order).”
Self-citationThe feasibility of artificial consciousness through the lens of neuroscience
Aru, J., Larkum, M. E., & Shine, J. M. (2023) · Trends in Neurosciences · doi:10.1016/j.tins.2023.09.009 · Cited by 98
Self-citationIntelligent nanophotonics: when machine learning sheds light
Wu, N., Sun, Y., Hu, J., Yang, C., Bai, Z., Wang, F. X., Cui, X., He, S., Li, Y., Zhang, C., et al. (2025) · eLight · doi:10.1186/s43593-025-00085-x · Cited by 91
Tononi, G., Boly, M., & Cirelli, C. (2024) · Neuron · doi:10.1016/j.neuron.2024.04.011 · Cited by 70
Brain-inspired artificial intelligence research: A review
Wang, G., Bao, H., Liu, Q., Zhou, T., Wu, S., Huang, T., Yu, Z., Lu, C., Gong, Y., Zhang, Z., et al. (2024) · Science China Technological Sciences · doi:10.1007/s11431-024-2732-9 · Cited by 43
Neuroethics and AI ethics: a proposal for collaboration
Salles, A. L. F., & Farisco, M. (2024) · BMC Neuroscience · doi:10.1186/s12868-024-00888-7 · Cited by 79
Artificial Intelligence and the Internal Processes of Creativity
Aru, J. (2025) · The Journal of Creative Behavior · doi:10.1002/jocb.1530 · Cited by 27
Self-citationBeyond Mechanism—Extending Our Concepts of Causation in Neuroscience
Potter, H. D., & Mitchell, K. J. (2025) · European Journal of Neuroscience · doi:10.1111/ejn.70064 · Cited by 25
Yan, H., Wang, L., Ma, K., & Zhong, Y. (2024) · IEEE Computer Society Conference on Computer Vision and Pattern Recognition/Proceedings - IEEE Computer Society Conference on Computer Vision and Pattern Recognition/Proceedings · doi:10.1109/cvpr52733.2024.02234 · Cited by 24
Predictive coding with spiking neurons and feedforward gist signaling
Lee, K., Dora, S., Mejias, J. F., Bohte, S. M., & Pennartz, C. M. A. (2024) · Frontiers in Computational Neuroscience · doi:10.3389/fncom.2024.1338280 · Cited by 14
“Such involvement of subcortical pathways is in line with the brain not being strictly hierarchical (Suzuki et al., 2023).”
BackgroundSelf-citationMultimodal mismatch responses in mouse auditory cortex
Sołyga, M., & Keller, G. B. (2025) · eLife · doi:10.7554/elife.95398.4 · Cited by 12
Imagination vs. routines: festive time, weekly time, and the predictive brain
Bortolotti, A., Conti, A., Romagnoli, A., & Sacco, P. L. (2024) · Frontiers in Human Neuroscience · doi:10.3389/fnhum.2024.1357354 · Cited by 13
Temporal prediction captures key differences between spiking excitatory and inhibitory V1 neuronsPreprint
Taylor, L., Zenke, F., King, A. J., & Harper, N. S. (2024) · bioRxiv (Cold Spring Harbor Laboratory) · doi:10.1101/2024.05.12.593763 · Cited by 10
Phenomenal interface theory: a model for basal consciousness
Klein, C., & Barron, A. B. (2025) · Philosophical Transactions of the Royal Society B: Biological Sciences · doi:10.1098/rstb.2024.0301 · Cited by 8
“In the typical mammalian brain, however, recurrent loops between the subcortex and regions of the cortex are a vital and almost universal feature of processing [85].”
Qi, X., Li, X., Lou, Y., Li, Y., Wang, G., Tang, K., & Zhao, J. P. (2024) · IEEE Journal of Solid-State Circuits · doi:10.1109/jssc.2024.3402208 · Cited by 8
Confidence and second-order errors in cortical circuits
Granier, A., Petrovici, M. A., Senn, W., & Wilmes, K. A. (2024) · PNAS Nexus · doi:10.1093/pnasnexus/pgae404 · Cited by 13
Yaroğlu, A. C. (2024) · Kybernetes · doi:10.1108/k-09-2024-2379 · Cited by 10
A compressed hierarchy for visual form processing in the tree shrew
Lanfranchi, F. F., Wekselblatt, J. B., Wagenaar, D. A., & Tsao, D. Y. (2025) · Nature · doi:10.1038/s41586-025-09441-w · Cited by 9
Prefrontal Dopamine in Flexible Adaptation to Environmental Changes: A Game for Two Players
Latagliata, E. C., Orsini, C., Cabib, S., Biagioni, F., Fornai, F., & Puglisi‐Allegra, S. (2023) · Biomedicines · doi:10.3390/biomedicines11123189 · Cited by 9
McKinnon, C., Mo, C., & Sherman, S. M. (2025) · Journal of Neuroscience · doi:10.1523/jneurosci.0002-25.2025 · Cited by 6
Brucklacher, M., Pezzulo, G., Mannella, F., Galati, G., & Pennartz, C. M. A. (2025) · Neural Networks · doi:10.1016/j.neunet.2024.106716 · Cited by 5
Self-citationA model of thalamo-cortical interaction for incremental binding in mental contour-tracing
Schmid, D., & Heiko Neumann, B. (2025) · PLoS Computational Biology · doi:10.1371/journal.pcbi.1012835 · Cited by 7
Cai, G., Chen, Y., Yang, B., Yang, Y., Ma, T., & Wang, Y. (2025) · Neural Networks · doi:10.1016/j.neunet.2025.107795 · Cited by 4
Multimodal mismatch responses in mouse auditory cortexPreprint
Sołyga, M., & Keller, G. B. (2024) · eLife · doi:10.7554/elife.95398 · Cited by 8
Wei, Z., Jin, F., Li, T., Qian, L., Ma, J., Liu, F., Zheng, W., Wang, Y., Zhang, S., You, Y. Z., et al. (2026) · Nature Communications · doi:10.1038/s41467-025-67723-3 · Cited by 3
Neuromorphic engineering: Artificial brains for artificial intelligence
Leugering, J. (2024) · Annals of the New York Academy of Sciences · doi:10.1111/nyas.15256 · Cited by 4
Network structure governs Drosophila brain functionality
Zhang, X., Yang, P., Feng, J., Wen, K., Yan, G., Luo, Q., Lin, W., & Lu, X. B. (2025) · Fundamental Research · doi:10.1016/j.fmre.2025.01.017 · Cited by 3
From hippocampal neurons to broad spiking neural networks
Wang, Y., Zuo, Y., Chen, D., Tu, W., Zomaya, A. Y., & Li, X. (2025) · Neurocomputing · doi:10.1016/j.neucom.2025.130547 · Cited by 3
Neuroscience-inspired continuous learning: a sustainable approach to AI energy challengePreprint
Xiao, D. (2023) · doi:10.31219/osf.io/twn9q · Cited by 3
Thalamo-Cortical Interaction for Incremental Binding in Mental Contour-TracingPreprint
Schmid, D., & Heiko Neumann, B. (2023) · bioRxiv (Cold Spring Harbor Laboratory) · doi:10.1101/2023.12.20.572705 · Cited by 3
A comparison between humans and AI at recognizing objects in unusual posesPreprint
Ollikka, N., Abbas, A., Perin, A., Kilpeläinen, M., & Deny, S. (2024) · PsyArXiv (OSF Preprints) · doi:10.31234/osf.io/z95t7 · Cited by 12
Evolutionary learning in neural networks by heterosynaptic plasticity
Bi, Z., Fu, R., Chen, G., Yang, D., Zhou, Y., & Tian, L. (2025) · iScience · doi:10.1016/j.isci.2025.112340 · Cited by 2
Gong, X., Chen, Y., Chen, C., Chen, C., & Zhang, X. (2025) · Food Bioscience · doi:10.1016/j.fbio.2025.107147 · Cited by 2
Du, Z., Chu, C., Shi, W., Luo, N., Lu, Y., Wang, H., Zhao, B., Xiong, H., Yang, Z., & Jiang, T. (2025) · Nature Communications · doi:10.1038/s41467-025-63204-9 · Cited by 2
Terlau, J., Martini, J., & Helfrich, R. F. (2025) · Neuron · doi:10.1016/j.neuron.2025.10.015 · Cited by 2
Shallow Brain Residual Network for Classifying UAVs and Birds in ISAC Base Stations
Sun, M., Yin, H., Wang, X., GuangXi, Z. H. U., & Debbah, M. (2025) · IEEE Vehicular Technology Conference · doi:10.1109/vtc2025-fall65116.2025.11309947 · Cited by 2
Eryeşil, K., & Öztürk, M. (2024) · Özgür Yayınları eBooks · doi:10.58830/ozgur.pub551.c2250 · Cited by 3
Multimodal mismatch responses in mouse auditory cortexPreprint
Sołyga, M., & Keller, G. B. (2024) · eLife · doi:10.7554/elife.95398.1 · Cited by 2
Multimodal mismatch responses in mouse auditory cortexPreprint
Sołyga, M., & Keller, G. B. (2024) · eLife · doi:10.7554/elife.95398.2 · Cited by 2
DeKraker, J., Eichert, N., Hong, S.-J., Evans, A. C., & Bernhardt, B. C. (2026) · Neuroscience & Biobehavioral Reviews · doi:10.1016/j.neubiorev.2026.106890 · Cited by 1
Sak, H. (2024) · Özgür Yayınları eBooks · doi:10.58830/ozgur.pub551.c2244 · Cited by 1
Li, J., & Wei, H. (2025) · IEEE Transactions on Image Processing · doi:10.1109/tip.2025.3562054 · Cited by 1
Wahidi, M., Rizk, A., & Imad, R. (2025) · IEEE Access · doi:10.1109/access.2025.3589514 · Cited by 1
Dynamic Reservoir Computing with Physical Neuromorphic Networks
Xu, Y., Gottwald, G. A., & Kuncic, Z. (2025) · International Joint Conference on Neural Networks (IJCNN) · doi:10.1109/ijcnn64981.2025.11229247 · Cited by 1
Control of representation updating by higher-order thalamus enables history-based decision-making
Hosford, P. S., Mei, H., Tagomori, H., Hayde, C., Aal, M. S. A. A., Tagomori, H., Miho, N., & Schmitt, L. I. (2026) · Neuron · doi:10.1016/j.neuron.2026.04.045 · Cited by 1
Fan, X., Chao, C., Zhang, L., Li, H., ZHAO, Y., Ding, Y., Bu, F., Zhou, W. Y., Liang, D., Chen, Y., et al. (2026) · Advanced Materials · doi:10.1002/adma.73590 · Cited by 1
Chu, Y., Li, G., Hou, S., Zhou, C., & Fei, J. (2026) · IEEE Internet of Things Journal · doi:10.1109/jiot.2026.3701685 · Cited by 1
The prelimbic cortex arbitrates memory-guided conflict resolution
Caban, A. J., Surets, M., Chien, C., Kaufmann, M., Park, S., & Ramirez, S. (2026) · Current Biology · doi:10.1016/j.cub.2026.06.047 · Cited by 2
Loops all the way up: Recurrency as an implementation-first primitive for consciousness
Zheng, Z., Chis-Ciure, R., Waade, P. T., Eiserbeck, A., Aru, J., Andrillon, T., Jarraya, B., Melloni, L., Northoff, G., Rosas, F. E., et al. (2026) · Physics of Life Reviews · doi:10.1016/j.plrev.2026.07.006 · Cited by 2
Self-citation‘Backpropagation and the brain’ realized in cortical error neuron microcircuits
Max, K., Jaras, I., Granier, A., Wilmes, K. A., & Petrovici, M. A. (2026) · PLoS Computational Biology · doi:10.1371/journal.pcbi.1014164 · Cited by 3
Switching Patterns of Cortical–Subcortical Interaction in the Human Brain
Nazzi, A., Favaretto, C., Vallesi, A., Corbetta, M., & Allegra, M. (2025) · Journal of Neuroscience · doi:10.1523/jneurosci.1855-24.2025 · Cited by 1
Lee, K., Baracco, L., Pennartz, C. M. A., Mototaka Suzuki, A., & Mejías, J. F. (2026) · Current Research in Neurobiology · doi:10.1016/j.crneur.2026.100155 · Cited by 1
Self-citationPredictive processing in neuroscience, computational modeling and psychology
Brucklacher, M., Lee, K., Moreni, G., Mejías, J. F., Bohté, S. M., & Pennartz, C. M. A. (2024) · Elsevier eBooks · doi:10.1016/b978-0-12-820480-1.00201-1 · Cited by 2
Self-citationMultimodal mismatch responses in mouse auditory cortexPreprint
Sołyga, M., & Keller, G. B. (2025) · eLife · doi:10.7554/elife.95398.3 · Cited by 1
The superior colliculus arbitrates whole-brain dynamics for unconscious visual insightPreprint
Murata, T., Kunishige, K., Seno, S., Ohzawa, I., Kaneko, K., Yanagida, T., Haruno, M., & Hosoda, K. (2025) · bioRxiv (Cold Spring Harbor Laboratory) · doi:10.1101/2025.04.09.648050 · Cited by 1
Behavioral hierarchy without a hierarchical brainPreprint
Han, Y., Liu, Q., Liu, S. Q., Cheng, H., & Wei, P. (2026) · bioRxiv (Cold Spring Harbor Laboratory) · doi:10.64898/2026.03.01.708904 · Cited by 1
Shallow Learning vs. Deep Learning in Finance, Marketing, and e-Commerce
Mageed, I. A., Bhat, A. H., & Edalatpanah, S. A. (2024) · The Springer series in applied machine learning · doi:10.1007/978-3-031-69499-8_3
Shallow Learning vs. Deep Learning in Social Applications
Mageed, I. A., Bhat, A. H., & Alja’am, J. M. (2024) · The Springer series in applied machine learning · doi:10.1007/978-3-031-69499-8_4
Prefrontal orchestration: a cortical network for rodent motor inhibitionPreprint
Jäckel, Z., Schwaderlapp, N., Adžemović, A., Steenbergen, F., Hardung, S., Fuchs, K., Leibold, C., Zaitsev, M., & Diester, I. (2024) · bioRxiv (Cold Spring Harbor Laboratory) · doi:10.1101/2024.10.16.618207
Günümüz(de) Liderlik Araştırmaları
Özkanan, A. (2024) · Özgür Yayınları eBooks · doi:10.58830/ozgur.pub551.c2241
Makyavelist Liderlik: Güç, Korku ve Manipülasyon
Güngör, B. (2024) · Özgür Yayınları eBooks · doi:10.58830/ozgur.pub551.c2248 · Cited by 2
Liderlik Dinamikleri Üzerine Araştırmalar II
Durmuş, G., Özkanan, A., Mutlu, M. D., Aytekin, M., Çelebi, B., Sak, H., Yaroğlu, A. C., Çakıroğlu, N., Uzun, A., Oğuzhan, Y. S., et al. (2024) · Özgür Yayınları eBooks · doi:10.58830/ozgur.pub551
Oğuzhan, Y. S. (2024) · Özgür Yayınları eBooks · doi:10.58830/ozgur.pub551.c2247 · Cited by 1
Yaroğlu, A. C. (2024) · Özgür Yayınları eBooks · doi:10.58830/ozgur.pub551.c2245
Cortical control of innate behavior from subcortical demonstrationPreprint
Keller, J. A., Kwak, I. S., Stark, A. K., Pachitariu, M., Branson, K. M., & Dudman, J. T. (2025) · bioRxiv (Cold Spring Harbor Laboratory) · doi:10.1101/2025.02.12.637930 · Cited by 1
A Hardware-Efficient Spiking Predictive Coding Architecture with Online STDP LearningPreprint
Kim, J.-G., Seo, D., Yeo, I., & Lee, B. (2025) · SSRN Electronic Journal · doi:10.2139/ssrn.5699160
Study on the Swimming Control of Bionic Robotic Fish Based on the Shallow Brain Model
Ma, H., Wang, M., Sun, H., Wang, C., Wu, Z., & Gao, H. (2025) · Communications in computer and information science · doi:10.1007/978-981-95-1103-7_23
Introduction to MRI Neuroanatomy
Duncan, J. S., & Giampiccolo, D. (2025) · Springer eBooks · doi:10.1007/978-3-031-76696-1_1
Brain-Inspired Broad Learning Framework: Spiking Neural Network for Enhanced EEG Prediction
Wang, Y., Zuo, Y., Chen, D., Tu, W., Chen, J., Gao, T., & Xiaoli Li, A. (2026) · IEEE Transactions on Emerging Topics in Computational Intelligence · doi:10.1109/tetci.2025.3647698
Li, J., Liu, X. Y., & Song, J. (2026) · IEEE Access · doi:10.1109/access.2026.3683697
Siegl, M., Haslgrübler, M., & Ferscha, A. (2026) · Communications in computer and information science · doi:10.1007/978-3-032-19099-4_25
Kim, J.-G., & Lee, B. (2026) · Applied Sciences · doi:10.3390/app16104896
A biological vision inspired framework for machine perception of illusory contours
Zhang, X., Yang, K.-F., Zhang, X.-S., You, H.-Z., Yan, H.-M., & Li, Y.-J. (2026) · Pattern Recognition · doi:10.1016/j.patcog.2026.114185
Pennartz, C. M. A., & Engeser, M. (2026) · Consciousness and Cognition · doi:10.1016/j.concog.2026.104077
Self-citationCross-modal invariants as the foundational basis of emerging brain circuitry representations
Ramasamy, A., & Jörntell, H. (2026) · Physics of Life Reviews · doi:10.1016/j.plrev.2026.06.010 · Cited by 2
Cellular mechanisms of serotonergic psychedelics - apical hypercontextualisation
Kaup, K., Jiménez, J. H., & Aru, J. (2026) · Neuroscience & Biobehavioral Reviews · doi:10.1016/j.neubiorev.2026.106876
Self-citationNeural selection in corticothalamocortical loops: Heterogeneity as a computational resource
Munn, B., Müller, E. J., Whyte, C. J., Brouwer, T., John, Y. J., Aru, J., Mototaka Suzuki, A., Saalmann, Y. B., Larkum, M. E., & Shine, J. M. (2026) · Progress in Neurobiology · doi:10.1016/j.pneurobio.2026.102945 · Cited by 1
Self-citationDorsomedial striatal neurons encode visual features
Perez-Becerra, J., Velazquez-Contreras, R., Téllez, L. A., & Carrillo‐Reid, L. (2026) · Cell Reports · doi:10.1016/j.celrep.2026.117958
Djebbara, Z., & Bogotá, J. D. (2026) · Neuroscience of Consciousness · doi:10.1093/nc/niaf050
Metadata
- Authors
- Suzuki, M., Pennartz, C. M. A., & Aru, J.
- Affiliations
- University of Amsterdam (NL)
- University of Tartu (EE)
- Corresponding author
- Aru, Jaan
- Venue
- Nature Reviews Neuroscience
- Published
- 2023-10-27
- Volume
- 24
- Issue
- 12
- Pages
- 778-791
- ISSN
- 1471-003X (ISSN-L)
- 1471-0048
- OpenAlex
- W4387996733
- Cited by
- 79
- References
- 224
- Citations by year
- 2026: 21 · 2025: 26 · 2024: 28 · 2023: 4
- FWCI
- 2.97
- Type
- journal article
- Language
- en
- Open access
- green
- handle.uba.uva.nl
- Licence
- Locations
- Nature reviews. Neuroscience (publishedVersion)
- PubMed (publishedVersion)
- UvA-DARE (University of Amsterdam) (publishedVersion, other-oa)
- Funders
- Brain Science Foundation
- European Commission (HORIZON2020, 945539)
- Eesti Teadusagentuur (PSG728)
- Topics
- Life Sciences › Neuroscience › Cognitive Neuroscience › Neural dynamics and brain function
- Life Sciences › Neuroscience › Cognitive Neuroscience › Memory and Neural Mechanisms
- Physical Sciences › Computer Science › Artificial Intelligence › Neural Networks and Applications
- Keywords
- thalamocortical loop, cortical microcircuits, predictive coding, deep learning architectures, neural network architecture, massively parallel computing
- MeSH terms
- Animals, Brain, Humans, Mammals, Neural Networks, Computer
About these checks
Registry, retraction, Cites and statistics results are computed, not judged, with one exception: a printed reference that no index lists is read once by a model before it counts as a citation error. Claims are extracted by a model and kept only when their quote appears verbatim in the text; each cited source is then checked, with a model’s help, for whether it says what the claim relies on. Those verdicts can be wrong — read the claim’s quote and the cited source before relying on one.
Statistics and arithmetic are recomputed, not judged.
Confidence bands are shown only for fields calibrated on our annotated test set. None is calibrated yet, so no band is shown.
What counts as a claim ›How we measure confidence ›How we check this ›
Are you an author, or do you see a problem with this page? Request a review