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Cited by 668

The DOI matches its Crossref record; 5 of the 5 references it cites were found in a registry or an index.

The STRING database in 2025: protein networks with directionality of regulation

Open access
Szklarczyk, Damian, Nastou, Katerina, Koutrouli, Mikaela, Kirsch, Rebecca, Mehryary, Farrokh, Hachilif, Radja, Hu, Dewei, Peluso, Matteo E, Huang, Qingyao, Fang, Tao, Doncheva, Nadezhda T, Pyysalo, Sampo, Bork, Peer, Jensen, Lars J, von Mering, Christian·2025·Nucleic Acids Research

Abstract

Proteins cooperate, regulate and bind each other to achieve their functions. Understanding the complex network of their interactions is essential for a systems-level description of cellular processes. The STRING database compiles, scores and integrates protein–protein association information drawn from experimental assays, computational predictions and prior knowledge. Its goal is to create comprehensive and objective global networks that encompass both physical and functional interactions. Additionally, STRING provides supplementary tools such as network clustering and pathway enrichment analysis. The latest version, STRING 12.5, introduces a new ‘regulatory network’, for which it gathers evidence on the type and directionality of interactions using curated pathway databases and a fine-tuned language model parsing the literature. This update enables users to visualize and access three distinct network types—functional, physical and regulatory—separately, each applicable to distinct research needs. In addition, the pathway enrichment detection functionality has been updated, with better false discovery rate corrections, redundancy filtering and improved visual displays. The resource now also offers improved annotations of clustered networks and provides users with downloadable network embeddings, which facilitate the use of STRING networks in machine learning and allow cross-species transfer of protein information. The STRING database is available online at https://string-db.org/.

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Crossref · 10.1093/nar/gkae1113

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Metadata

Authors
Szklarczyk, D., Nastou, K., Koutrouli, M., Kirsch, R., Mehryary, F., Hachilif, R., Hu, D., Peluso, M. E., Huang, Q., Fang, T., Doncheva, N. T., Pyysalo, S., Bork, P., Jensen, L. J., & von Mering, C.
Venue
Nucleic Acids Research
Published
2024-11-18
Volume
53
Issue
D1
Pages
D730-D737
ISSN
  • 0305-1048 (ISSN-L)
  • 1362-4962
OpenAlex
W4404511387
Cited by
668
References
5
FWCI
77.61
Type
journal article
Language
en
Open access
gold

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