Forest Succession in Relation to River Terrace Development in Olympic National Park, Washington

Abstract

The floodway zone of the Hoh River exhibits four terrace levels of different ages, formed by erosional activity of the river on valley fills. The vegetation in this valley is in a long—term seral sequence as shown by the zonal pattern in relation to aging and development of these land surfaces. Succession starts on gravel bars, which are dominated by Alnus rubra and Salix scouleriana. The following sequential forest communities, and associated ages of land surfaces, are found: Alnus rubra on alder flats (80—100 yr); Picea sitchensis—Acer macrophyllum—Populus trichocarpa on first terraces (400 yr); Picea sitchensis—Tsuga heterophylla on second terraces (750 yr); and Tsuga heterophylla on third terraces. The latter represents the climax community for the river terrace sere, and it occurs on surfaces exposed by retreating Pleistocene alpine glaciers. The first three terraces are derived from Neoglacial alluvial fills. There is a strong correlation among zonation patterns, forest succession, age of terraces, soil moisture, and soil profile development. Available soil moisture is an important factor governing the zonal sequence. The younger land surfaces are significantly drier than the older terraces. Plants on alder flats and first terraces must withstand greater moisture stress than those of second and third terraces. As the land surface ages, the soil profile develops; deeper, more mature soils are found away from the river. The term "Olympic rain forest" is inappropriately applied to this vegetation; "temperate moist coniferous forest" is more appropriate not only for forests in the Hoh Valley, but also for the rest of the Olympic Mountains and vegetation along the northern Pacific coast.