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Forest stand structure and functioning: Current knowledge and future challenges
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“Regulating stands into a more complex vertical structure will lead to faster growth and greater C sequestration in forests because multilayered canopies will occupy a range of light environments, resulting in high light acquisition and light-use efficiency.180,181”
From Technologies and perspectives for achieving carbon neutrality (2021) · cites this paper as [180]
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Rebollo, P., Cruz‐Alonso, V., Ruiz‐Benito, P., Triviño, M., Astigarraga, J., González‐Díaz, P., Zavala, M. A., & Andivia, E. (2025) · Journal of Ecology · doi:10.1111/1365-2745.70183 · Cited by 2
“This allowed us to assess whether struc - tural diversity contributed to above-ground productivity through po - tential complementarity effects mediated by size differences among individuals (Ali, 2019).”
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“The asymmetric competition for light, for instance, is noted in a forest structure meta-study as a key negative influence on overall productivity [9].”
BackgroundPraças urbanas como potenciais ferramentas na neutralização de carbono
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“The insignificance of the connections between homogenous forest stands and soil diversity signals that monocoenoses do not importantly increase the topsoil diversity (Ali, 2019).”
BackgroundYin, P., Zhai, K., Zhao, X., Li, R., Fernández‐Alonso, M. J., Wang, J., Yang, H., Zhang, X., Berg, B., Wang, S., et al. (2025) · Journal of Ecology · doi:10.1111/1365-2745.70141 · Cited by 1
“…we estimated, for each plot, the basal area (BA) using diameter at breast height (DBH) of all woody plants as a proxy of vegetation biomass, species richness (SR) as a proxy of diversity and the co - efficient of variation of DBH (CVD) as a proxy of forest structural diversity (Ali, 2019 ).”
Mashhadi, N., & Sánchez-Azofeifa, A. G. (2025) · International Journal of Applied Earth Observation and Geoinformation · doi:10.1016/j.jag.2025.104922 · Cited by 1
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“The absence of carbon-mortality trade-offs in natural forests suggests that mimicking their structural diversity could reduce vulnerabilities in planted systems (Ali et al. 2016, 2019).”
Jiang, S., Deng, X., Huang, Z., Xie, W., Zhu, W., Xiang, W., & Lu, X. (2026) · Trees · doi:10.1007/s00468-026-02742-1 · Cited by 1
“…of different stands, even though the effects were not significant ( p > 0.05), indicating that competition an important driver of complementarity (Ali 2019), stand structural diversity is related to stand biomass retention rate, which helps to predict forest growth and stand dynamics (Gough et…”
InfluentialExplaining the relationship between structural diversity, tree and stand growth
Pretzsch, H., del Rı́o, M., Ruíz‐Peinado, R., Heym, M., Bielak, K., Brazaitis, G., Granhus, A., Jansons, Ā., Holm, S., Löf, M., et al. (2026) · Forest Ecology and Management · doi:10.1016/j.foreco.2026.123699 · Cited by 1
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Comparing Two Novel LiDAR ‐Based Indices for Quantifying Forest Structural Complexity
Reuter, T., Seidel, S., & Seidel, D. (2026) · Ecology and Evolution · doi:10.1002/ece3.73605 · Cited by 1
“A recent study emphasizes the need for a unified methodology to assess forest structure in order to gain knowledge about the interrelations of forest structure with ecosystem functions (Ali 2019).”
Optimizing urban park ecological benefits: Linking 3D structure to function with LiDAR and GAMs
Zhao, Y., Liu, H., Li, H., Liu, C., Jiang, H., Dong, N., Ge, S., & Tian, G. (2026) · Ecological Informatics · doi:10.1016/j.ecoinf.2026.103858 · Cited by 1
Canopy structural complexity promotes ecosystem multifunctionality in a temperate forest
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“On a medium-to long-term scale, we recommend the creation of a municipal protected area covering the studied fragment.”
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“…relationships between stand structure and biodiversity (Atikah et al., 2021; Gebremichael et al., 2022; Penone et al., 2019; Storch et al., 2023), and between stand structure and ecosystem functions (Ali, 2019; Felipe-Lucia et al., 2018; Firn et al., 2007) are highly variable and context-specific.”
BackgroundLa jurisdicción constitucional y el modelo dual o paralelo
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Anwar, M., Hanif, A., Zhanwu, G. A. O., Rasheed, A., Shahzad, S., Haseeb, A., Gul, M., Al-Khayri, J. M., Al-Daej, M. I., Sattar, M. N., et al. (2023) · Notulae Botanicae Horti Agrobotanici Cluj-Napoca · doi:10.15835/nbha51413370
Title: Surface Runoff Simulation Using Swat Analysis in Upper Gillo Watershed, EthiopiaPreprint
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Bioindicators for Forest Area Condition: A Systematic Literature Review
Rahardjanto, A., & Husamah, H. (2024) · Jurnal Penelitian dan Pengkajian Ilmu Pendidikan e-Saintika · doi:10.36312/esaintika.v8i1.1793
“…presence of diverse tree species and other plants signifies ecosystem stability, shapes forest structure, and influences the ecological functioning of the forest (Ali, 2019; Bugmann & Seidl, 2022; Ikbal et al., 2023; Latt & Park, 2022; Nugroho et al., 2022; Pan et al., 2018; Y. Wang et al., 2023).”
BackgroundEffect of Thinning on Cryptomeria Japonica Tree Size Differentiation and Stand ProductivityPreprint
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Estoques aéreos de biomassa e carbono em remanescente de Caatinga no Estado de Sergipe, Brasil
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“Understory vegetation is a vital component of forest eco-systems, supporting essential ecosystem functions such as C storage, primary production and nutrient supply (Ali et al., 2019).”
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Zhao, X., Xin, Y., Xie, P., Wang, B., Lv, X., Wan, P., Wang, P., & Ma, H. (2025) · Environmental Management · doi:10.1007/s00267-025-02293-0
Pretzsch, H., del Rı́o, M., Ruíz‐Peinado, R., Heym, M., Bielak, K., Brazaitis, G., Granhus, A., Jansons, Ā., Holm, S., Löf, M., et al. (2025) · SSRN Electronic Journal · doi:10.2139/ssrn.5822827
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Additional file 1 of The relationships among structure variables of larch forests in China
Fang, W., Zhao, Q., Cai, Q., Eziz, A., Chen, G., Feng, Y., Zhang, H., Zhu, J., Ji, C., Tang, Z., et al. (2020) · Figshare · doi:10.6084/m9.figshare.13238588
Wang, Z., Qin, J., Yang, L., He, J., Hao, M., Liu, G., He, H., Zhao, X., von Gadow, K., Zhang, C., et al. (2026) · Journal of Environmental Management · doi:10.1016/j.jenvman.2026.128771
Zou, K., Liu, X., Gao, L., Yang, L., Qin, J., Xu, Q., & He, P. (2026) · Forest Ecology and Management · doi:10.1016/j.foreco.2026.123587
Wan, D., Wang, Q., Wang, H., Chen, Q., & Wang, Y. (2026) · Forest Science · doi:10.1007/s44391-025-00055-3
Phylogeny sustains the functional diversity of temperate forests under environmental changePreprint
Gu, Y., Zhang, J., & Shi-jie, H. (2026) · SSRN Electronic Journal · doi:10.2139/ssrn.6222327
Pretzsch, H. (2026) · Forest Ecology and Management · doi:10.1016/j.foreco.2026.123627
Orou, H., Assèdé, E. S. P., Ponette, Q., & Biaou, S. S. H. (2026) · Frontiers in Forests and Global Change · doi:10.3389/ffgc.2026.1800379
Diameter class-based model for predicting forest growth dynamics and responses to thinning regimes
Lee, J.-A., Kwon, S. J., Noulèkoun, F., Kim, H., Lee, J.-M., Ku, M., Jo, H., & Son, Y. (2026) · Forest Ecology and Management · doi:10.1016/j.foreco.2026.123680
Explainable machine learning identifies key degradation factors in Robinia pseudoacacia shelterbelts
Ye, P., Lu, P., Xie, P., Wang, X., He, Y., Zhou, M., Jin, S., & Yan, D. (2026) · Restoration Ecology · doi:10.1111/rec.70395
“The positive association between the W and degradation indicates that a loss of spatial regularity intensi fi es resource competition in the nutrient-poor sandy substrate, destabilizing stand structure (Ali 2019; Wang et al. 2022).”
Khanal, S., Koirala, M., Shrestha, N., Gautam, P., Lama, F., Degen, A., Joshi, S., Sparrow, B. D., & Shang, Z. (2026) · Global Ecology and Conservation · doi:10.1016/j.gecco.2026.e04225
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Okullo, J. B. L., Sylvano, A., Nangendo, G., & Kalema, J. (2026) · International Journal of Forestry Research · doi:10.1155/ijfr/5524104
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Biyiha, E. J., Ebanga, A. P., Kpoumie, H. M., Yonga, G., Fobane, J. L., Chimi, C. D., Messado, K. L., Makak, R. N., Nguihi, L. M., & Zapfack, L. (2026) · Research Square · doi:10.21203/rs.3.rs-9367677/v1
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Factors determining the behaviour of dynamic self-thinning lines in Canadian interior spruce forests
West, P. W. (2026) · Trees · doi:10.1007/s00468-026-02787-2
“The processes become more complex in uneven-aged and / or mixed species forests (e.g. del Río et al. 2014, 2016; For-rester and Bauhaus 2016; Ali 2019; Trogisch et al. 2021; Acquah et al. 2025; Dong et al. 2025).”
Kweon, D., & Jo, J. (2026) · SSRN Electronic Journal · doi:10.2139/ssrn.6873736
Effects of stand structure on leaf biomass and productivity in Larix kaempferi plantation
Wang, C., Chen, D., Chunyu, G. E., & Tang, X. (2026) · Forest Ecology and Management · doi:10.1016/j.foreco.2026.123998
Wang, J., Lv, C., Yin, J., Chen, Y., Lv, S., & Wu, B. (2026) · Frontiers in Plant Science · doi:10.3389/fpls.2026.1840683
“Optimising stand structure enhances long-term productivity and economic bene fi ts, thereby supporting sustainable forest development (Huang et al., 2019; Ali, 2019; Tang et al., 2020; Von Gadow and Hui, 2002).”
Liu, C., He, H., Huang, L., Chen, G., Liu, Z., Dou, X., & Zhu, G. (2026) · Ecological Indicators · doi:10.1016/j.ecolind.2026.115193
Species Turnover and Environmental Filtering Structure Plant Communities in Moist Temperate Forests
Samad, M., Ahmad, N., Ahmad, A., Al‐Andal, A., Khan, A., Osmani, A. R., & Waheed, M. (2026) · Ecology and Evolution · doi:10.1002/ece3.74054
“Existing research often treats these components separately, limiting mechanistic understanding of forest assembly processes (Weiher et al. 2011; Ali 2019; Sanders and Frago 2024).”
Chen, M., Wang, C., Zhang, Z., Ma, M., Zou, W., & Peng, D. (2026) · Journal of Environmental Management · doi:10.1016/j.jenvman.2026.130607
Chaves, J. E., Rodríguez‐Souilla, J., Cellini, J. M., Lencinas, M. V., Peri, P. L., & Pastur, G. J. M. (2026) · Ecological Processes · doi:10.1186/s13717-026-00740-2
“Forest structure is associated with a variety of processes and functions, such as biomass accumulation (Ali 2019), carbon storage (Hardi-man et al. 2013), primary production and nutrient uptake (Bechtold et al. 2017).”
Shangguan, Z., Wang, Y., Qin, J., Liu, Z., Yu, H., Feng, T., Wang, C., Lang, T., & Wang, P. (2026) · Plant and Soil · doi:10.1007/s11104-026-08990-x
“Nevertheless, the overall higher spatial structural complexity—particularly in terms of canopy species mingling and intense interspecific competition observed in the secondary forest—is associated with greater community composition stability and enhanced ecosystem functions (Ali 2019).”
Yan, P., Gao, Q., Wu, Y., Xu, Y., Luo, J., Li, Y., Zhao, W., & Li, C. (2026) · SSRN Electronic Journal · doi:10.2139/ssrn.7353762
Louzada, R. O., de Vasconcelos, D. C., Bozza, A. N., Chaib, J. G., & Tavares, R. M. B. (2026) · Forest Ecology and Management · doi:10.1016/j.foreco.2026.124190
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Li, S., Wei, T., Huang, J., Zhu, Q., & Fan, D. (2026) · Plant and Soil · doi:10.1007/s11104-026-09060-y
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Declining forest structural heterogeneity triggers multi-trophic species diversity lossPreprint
Wang, T., Dong, L., & Liu, Z. (2026) · SSRN Electronic Journal · doi:10.2139/ssrn.7450877
Integrating stand and environmental factors to enhance aboveground carbon stock estimation
Zhou, X., Guo, J., Sharma, R. P., & Guan, F. (2026) · Trees · doi:10.1007/s00468-026-02810-6
“To better reflect environ - mental effects (Ali 2019; Grace et al. 2016), the selection threshold was relaxed to |r| > 0.1 and VIF < 5, enabling the development of models M1-M5 with various combinations of predictors.”
InfluentialKim, T.-R., Lee, M.-K., Lim, J.-H., Lee, S.-T., Chung, S., & Lee, C. (2026) · Environmental Technology & Innovation · doi:10.1016/j.eti.2026.105234
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Effects of structural diversity on the productivity of oak forests in the Qinling Mountains, China
Zhang, G., Liu, Q., Wang, Y., Jiang, Y., von Gadow, K., & Yu, F. (2026) · European Journal of Forest Research · doi:10.1007/s10342-026-01957-8
“This effect is consis - tent with findings across forest biomes (Ali et al. 2016 ; Ali 2019 ; Aponte et al. 2020 ), where it is primarily mediated by enhanced light capture efficiency in mixed-aged stands (Ali et al. 2020b ).”
InfluentialSimonelli, F. G., Cocozza, C., Rosellini, I., Traversari, S., Campagnaro, T., Vítková, M., & Sitzia, T. (2026) · European Journal of Forest Research · doi:10.1007/s10342-026-01955-w
“…attributes such as stem density, tree diameter and height distribution, basal area, stand volume and canopy development are associated with ecosystem functioning, including biomass and carbon storage, light and microclimatic regulation, and habitat pro-vision (McElhinny et al. 2005; Ali 2019).”
Zhai, Y., Wang, L., Rahman, M. A., Yao, Y., Jia, J., Hao, S., Li, R., Zhang, X., Chen, Y., Chang, B., et al. (2026) · Building and Environment · doi:10.1016/j.buildenv.2026.115394
Drivers of aboveground biomass along riparian gradients in desert Populus euphratica forests
Liu, H., Wang, J., Yang, Z., Zhu, L., Zhai, J., & Li, Z. (2026) · Forest Ecosystems · doi:10.1016/j.fecs.2026.100548
Competition modulates growth in Pinus canariensis plantations: Implications for precision thinning
Navarro-Cerrillo, R. M., Baietto, A., González, A. C., Valeriano, C., Cedrés, E. P., & Camarero, J. J. (2026) · Forest Ecosystems · doi:10.1016/j.fecs.2026.100549
Lisner, A., Lepš, J., Novotný, V., Butterill, P. T., Houdková, L., Molem, K., Philip, J., Weiblen, G. D., & Fibich, P. (2026) · Oikos · doi:10.1002/oik.12452
Metadata
- Authors
- Ali, A.
- Affiliations
- South China Normal University (CN)
- Corresponding author
- Ali, Arshad
- ORCID
- Ali, Arshad
- Venue
- Ecological Indicators
- Publisher
- Elsevier BV
- Published
- 2019-03
- Volume
- 98
- Pages
- 665-677
- ISSN
- 1470-160X (ISSN-L)
- OpenAlex
- W2900494555
- Cited by
- 296
- References
- 62
- Citations by year
- 2026: 55 · 2025: 70 · 2024: 50 · 2023: 47 · 2022: 28 · 2021: 25 · 2020: 13 · 2019: 7
- FWCI
- 8.83
- Type
- journal article
- Language
- en
- Open access
- closed
- Locations
- Ecological Indicators (publishedVersion)
- Funders
- Guangdong Provincial Government
- South China Normal University
- Topics
- Life Sciences › Agricultural and Biological Sciences › Forestry › Forest ecology and management
- Physical Sciences › Environmental Science › Nature and Landscape Conservation › Ecology and Vegetation Dynamics Studies
- Life Sciences › Agricultural and Biological Sciences › Forestry › Remote Sensing and LiDAR Applications
- Keywords
- forest stand structure, forest functioning, species diversity, aboveground biomass, carbon storage, forest productivity, niche complementarity, plant-plant interactions, light competition, competitive exclusion, agroforests, plantations, tropical forests, temperate forests, boreal forest
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