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The Effects of Developer Dynamics on Fitness in an Evolutionary Ecosystem Model of the App Store
Abstract
Natural ecosystems exhibit complex dynamics of interacting species. Man-made ecosystems exhibit similar dynamics and, in the case of mobile app stores, can be said to perform optimization as developers seek to maximize app downloads. This work aims to understand stability and instability within app store dynamics and how it affects fitness. The investigation is carried out with AppEco, a model of the iOS App Store, which was extended for this paper and updated to model the store from 2008 to 2014. AppEco models apps containing features, developers who build the apps, users who download apps according to their preferences, and an app store that presents apps to the users. It also models developers who use commonly observed strategies to build their apps: innovator, milker, optimizer, copycat, and flexible (the ability to choose any strategy). Results show that despite the success of the copycat strategy, there is a clear stable state for low proportion of copycats in developer populations, mirroring results in theoretical biology for producer-scrounger systems. The results also show that the best fitness is achieved when the evolutionary optimizer (as producer) and copycat (as scrounger) strategies coexist together in stable proportions.
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“When a unique new product or feature enters the ecosystem, copies of it in somewhat altered form follow (e.g., when Flappy Bird was released, many clones appeared in the appstore in the following months [74]).”
From The Impact of Generative AI on Creativity in Software Development: A Research Agenda (2025) · cites this paper as [74]
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Cited by
Sorted by citations
The Impact of Generative AI on Creativity in Software Development: A Research Agenda
Jackson, V., Vasilescu, B., Russo, D., Ralph, P., Prikladnicki, R., Izadi, M., D’Angelo, S., Inman, S., Andrade, A., & van der Hoek, A. (2025) · ACM Transactions on Software Engineering and Methodology · doi:10.1145/3708523 · Cited by 23
Metadata
- Authors
- Lim, S. L., Bentley, P. J., & Ishikawa, F.
- Affiliations
- University College London (GB)
- National Institute of Informatics (JP)
- Venue
- IEEE Transactions on Evolutionary Computation
- Published
- 2016-08
- Volume
- 20
- Issue
- 4
- Pages
- 529-545
- ISSN
- 1089-778X (ISSN-L)
- 1941-0026
- OpenAlex
- W2345110451
- Cited by
- 13
- References
- 42
- Citations by year
- 2026: 1 · 2024: 1 · 2023: 3 · 2022: 3 · 2021: 1 · 2020: 1 · 2018: 1 · 2017: 1 · 2016: 1
- FWCI
- 3.32
- Type
- journal article
- Language
- en
- Open access
- closed
- Locations
- IEEE Transactions on Evolutionary Computation (publishedVersion)
- Topics
- Social Sciences › Sociology and Political Science › Evolutionary Game Theory and Cooperation
- Social Sciences › Business, Management and Accounting › Strategy and Management › Digital Platforms and Economics
- Life Sciences › Biochemistry, Genetics and Molecular Biology › Genetics › Evolution and Genetic Dynamics
- Keywords
- iOS App Store, mobile app stores, developer strategies, milkers, evolutionary stability, coexistence, fitness, ecosystem dynamics
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