The DOI matches its Crossref record; 37 of the 37 references it cites were found in a registry or an index.
The tenth skeletal specimen of Archaeopteryx
Open accessAbstract
We describe the tenth skeletal specimen of the Upper Jurassic Archaeopterygidae. The almost complete and well-preserved skeleton is assigned to Archaeopteryx siemensii Dames, 1897 and provides significant new information on the osteology of the Archaeopterygidae. As is evident from the new specimen, the palatine of Archaeopteryx was tetraradiate as in non-avian theropods, and not triradiate as in other avians. Also with respect to the position of the ectopterygoid, the data obtained from the new specimen lead to a revision of a previous reconstruction of the palate of Archaeopteryx. The morphology of the coracoid and that of the proximal tarsals is, for the first time, clearly visible in the new specimen. The new specimen demonstrates the presence of a hyperextendible second toe in Archaeopteryx. This feature is otherwise known only from the basal avian Rahonavis and deinonychosaurs (Dromaeosauridae and Troodontidae), and its presence in Archaeopteryx provides additional evidence for a close relationship between deinonychosaurs and avians. The new specimen also shows that the first toe of Archaeopteryx was not fully reversed but spread medially, supporting previous assumptions that Archaeopteryx was only facultatively arboreal. Finally, we comment on the taxonomic composition of the Archaeopterygidae and conclude that Archaeopteryx bavarica Wellnhofer, 1993 is likely to be a junior synonym of A. siemensii, and Wellnhoferia grandis Elżanowski, 2001 a junior synonym of A. lithographica von Meyer, 1861.
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“All non-avian theropods, and the first bird Archaeopteryx, lack a reversed hallux [20].”
From Assessing Arboreal Adaptations of Bird Antecedents: Testing the Ecological Setting of the Origin of the Avian Flight Stroke (2011) · cites this paper as [20]
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“This would be particularly acute in non-avian theropod like Microraptor, given the lack of a reversed hallux to aid in securing a purchase [19], [20].”
From Assessing Arboreal Adaptations of Bird Antecedents: Testing the Ecological Setting of the Origin of the Avian Flight Stroke (2011) · cites this paper as [20]
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Citations
Cites
Saurischian monophyly and the origin of birds
Gauthier, J. (1986) · The Academy · doi:10.5281/zenodo.16171435 · Cited by 1,294
Sereno, P. C. (1999) · Science · doi:10.1126/science.284.5423.2137 · Cited by 835
Four-winged dinosaurs from China
Xu, X., Zhou, Z., Wang, X., Kuang, X., Zhang, F., & Du, X. (2003) · Nature · doi:10.1038/nature01342 · Cited by 485
Archaeopteryx and the origin of birds
OSTROM, J. H. (1976) · Biological Journal of the Linnean Society · doi:10.1111/j.1095-8312.1976.tb00244.x · Cited by 361
(1999) · Bulletin of the American Museum of Natural History · Cited by 346
A dromaeosaurid dinosaur with a filamentous integument from the Yixian Formation of China
Xu, X., Wang, X.-L., & Wu, X.-C. (1999) · Nature · doi:10.1038/45769 · Cited by 314
The theropod ancestry of birds: new evidence from the late cretaceous of madagascar
Forster, C. A., Sampson, S. D., Chiappe, L. M., & Krause, D. W. (1998) · Science · doi:10.1126/science.279.5358.1915 · Cited by 314 · cited 6×
“This feature is otherwise known only from the basal avian Rahonavis and deinonychosaurs (Dromaeosauridae and Troodontidae), and its presence in Archaeopteryx provides additional evidence for a close relationship between deinonychosaurs and avians.”
BackgroundResultInfluentialA long-tailed, seed-eating bird from the Early Cretaceous of China
Zhou, Z., & Zhang, F. (2002) · Nature · doi:10.1038/nature00930 · Cited by 274
Olson, S. (2000) · The Auk · doi:10.1642/0004-8038(2000)117[0836:aasotc]2.0.co;2 · Cited by 259
Clarke, J. A. (2004) · Bulletin of the American Museum of Natural History · doi:10.1206/0003-0090(2004)286 · Cited by 248
Research works printed but not linked: 22
- 1990, Ornithomimosauria, 225
- 1993, Osteologia, Handbook of avian anatomy: Nomina Anatomica Avium. Publications of the Nuttall Ornithological Club, 23, 45
- 1999, A comparison of the jaw skeleton in theropods and birds, with a description of the palate in the Oviraptoridae, Smithsonian Contributions to Paleobiology, 89, 311
- 2002, Archaeopterygidae (Upper Jurassic of Germany), 129
- 2001, The function of the manus and forelimb of Deinonychus antirrhopus and its importance for the origin of avian flight, 301
- 1982, Zur Fossilisation der Federn des Urvogels (Berliner Exemplar). Wissenschaftliche Zeitschrift der Humboldt-Universität zu Berlin, Mathematisch-Naturwissenschaftliche Reihe, 31, 185
- 1985, The relationship of Archaeopteryx to other birds, 177
- 1991, Mesozoic birds and the origin of birds, 485
- 1997, Der achte Archaeopteryx, Fossilien, 3/97, 156
- 2002, Evolution of the perching foot in theropods, Journal of Vertebrate Paleontology, 22, 88A
- 1991, The question of the origin of birds, 467
- 1996, Ein nahezu vollständiges Skelett eines urtümlichen Vogels aus China, Natur und Museum, 126, 298
- 1992, A comparative study on the claws of Archaeopteryx, Los Angeles County Museum of Natural History, Contributions to Science, 36, 29
- 1985, Feathers and wings of Archaeopteryx, and the question of her flight ability, 251
- 2004, Kurznotiz: Nachweis von Überresten eines neuen Exemplars des Urvogels Archaeopteryx aus Solnhofen, Archaeopteryx, 22, 1
- 1987, Urvögel Archaeopterygiformes
- 1991, Archaeopteryx: quo vadis?, 485
- 1992, A new specimen of Archaeopteryx from the Solnhofen limestone, Los Angeles County Museum of Natural History, Science Series, 36, 3
- 1993, Das siebte Exemplar von Archaeopteryx aus den Solnhofener Schichten, Archaeopteryx, 11, 1
- 2005, Das neunte Archaeopteryx-Exemplar von Solnhofen, Archaeopteryx, 23, 3
- 2004, Das ‘Brustbein’ von Archaeopteryx bavarica Wellnhofer 1993–eine Revision, Archaeopteryx, 22, 3
- 2002, The debate on avian ancestry: phylogeny, function, and fossils, 3
Cited by
Sorted by citations
A Review of Dromaeosaurid Systematics and Paravian Phylogeny
Turner, A. H., Makovicky, P. J., & Norell, M. A. (2012) · Bulletin of the American Museum of Natural History · doi:10.1206/748.1 · Cited by 350
“, Velociraptor mongoliensis IGM 100/985, Archaeopteryx lithographica [Mayr et al., 2007], Mei long IVPP 12733).”
BackgroundAn integrative approach to understanding bird origins
Xu, X., Zhou, Z., Dudley, R., Mackem, S., Chuong, C.-M., Erickson, G. M., & Varricchio, D. J. (2014) · Science · doi:10.1126/science.1253293 · Cited by 319
“specialized second pedal digit as in deinonychosaurs (61); Jeholornis (Fig.”
BackgroundAn Archaeopteryx-like theropod from China and the origin of Avialae
Xu, X., You, H., Du, K., & Han, F. (2011) · Nature · doi:10.1038/nature10288 · Cited by 311
A Jurassic avialan dinosaur from China resolves the early phylogenetic history of birds
Godefroit, P., Cau, A., Dong-Yu, H., Escuillié, F., Wenhao, W., & Dyke, G. (2013) · Nature · doi:10.1038/nature12168 · Cited by 216
Benson, R., Butler, R. J., Carrano, M. T., & O’Connor, P. M. (2011) · Biological reviews/Biological reviews of the Cambridge Philosophical Society · doi:10.1111/j.1469-185x.2011.00190.x · Cited by 175
“Although few unambiguously pneumatic vertebrae are well preserved in Archaeopteryx, cervical vertebrae are unambiguously pneumatic, and middle-posterior dorsal and caudal vertebrae are definitely apneumatic.”
BackgroundInfluential
Metadata
- Authors
- MAYR, G., POHL, B., HARTMAN, S., & PETERS, D. S.
- Affiliations
- Senckenberg Research Institute and Natural History Museum Frankfurt/M (DE)
- ORCID
- HARTMAN, SCOTT
- Venue
- Zoological Journal of the Linnean Society
- Publisher
- Oxford University Press (OUP)
- Published
- 2007-01
- Volume
- 149
- Issue
- 1
- Pages
- 97-116
- ISSN
- 0024-4082 (ISSN-L)
- 1096-3642
- OpenAlex
- W2024552857
- Cited by
- 149
- References
- 37
- Citations by year
- 2026: 9 · 2025: 11 · 2024: 3 · 2023: 4 · 2022: 5 · 2021: 8 · 2020: 6 · 2019: 8 · 2018: 16 · 2017: 12 · 2016: 8 · 2015: 3 · 2014: 5 · 2013: 11 · 2012: 6
- FWCI
- 5.83
- Type
- journal article
- Language
- en
- Open access
- bronze
- doi.org
- Locations
- Zoological Journal of the Linnean Society (publishedVersion)
- Zenodo (CERN European Organization for Nuclear Research) (submittedVersion, cc-by)
- Topics
- Physical Sciences › Earth and Planetary Sciences › Paleontology › Paleontology and Evolutionary Biology
- Physical Sciences › Earth and Planetary Sciences › Paleontology › Evolution and Paleontology Studies
- Life Sciences › Agricultural and Biological Sciences › Ecology, Evolution, Behavior and Systematics › Fossil Insects in Amber
- Keywords
- osteology, ectopterygoid, coracoid process, tarsal bones, avian origin, arboreality, junior synonym, Late Jurassic
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