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New fossils from Jebel Irhoud, Morocco and the pan-African origin of Homo sapiens.
Open accessAbstract
Fossil evidence points to an African origin of Homo sapiens from a group called either H. heidelbergensis or H. rhodesiensis. However, the exact place and time of emergence of H. sapiens remain obscure because the fossil record is scarce and the chronological age of many key specimens remains uncertain. In particular, it is unclear whether the present day 'modern' morphology rapidly emerged approximately 200 thousand years ago (ka) among earlier representatives of H. sapiens or evolved gradually over the last 400 thousand years. Here we report newly discovered human fossils from Jebel Irhoud, Morocco, and interpret the affinities of the hominins from this site with other archaic and recent human groups. We identified a mosaic of features including facial, mandibular and dental morphology that aligns the Jebel Irhoud material with early or recent anatomically modern humans and more primitive neurocranial and endocranial morphology. In combination with an age of 315 ± 34 thousand years (as determined by thermoluminescence dating), this evidence makes Jebel Irhoud the oldest and richest African Middle Stone Age hominin site that documents early stages of the H. sapiens clade in which key features of modern morphology were established. Furthermore, it shows that the evolutionary processes behind the emergence of H. sapiens involved the whole African continent.
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“a recent study reported human remains with many modern features but archaic cranial morphology dated to about 315 ka ago (38), suggesting that not all human-specific traits arose at the same time.”
From An ancestral recombination graph of human, Neanderthal, and Denisovan genomes (2021) · cites this paper as [38]
The cited source says what the claim attributes to it.
In the citing paper
“All anatomically modern humans (AMHs) trace their origin to the African continent around 300 ka (Stringer 2016; Hublin et al. 2017)”
From Archaic Introgression Shaped Human Circadian Traits (2023) · cites this paper as Hublin et al. 2017
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In the citing paper
“The emergence of the earliest hominins possessing most anatomical traits associated with H. sapiens has been traced back to Jebel Irhoud, Morocco, around 300,000 years BP (Hublin et al., 2017).”
From The evolution of human skin pigmentation: A changing medley of vitamins, genetic variability, and UV radiation during human expansion (2023) · cites this paper as Hublin et al., 2017
In the citing paper
“The Jebel Irhoud specimens have short faces, small teeth, and reduced brow-ridges compared to pre-sapiens ancestors, making them the earliest human specimens to show features of the domestication syndrome.”
From Hypotheses for the Evolution of Reduced Reactive Aggression in the Context of Human Self-Domestication (2019) · cites this paper as Hublin et al., 2017
In the citing paper
“Currently the earliest record of a lineage of H. sapiens comes from around 315,000 years ago from Jebel Irhoud, Morocco (Hublin et al., 2017).”
From Hypotheses for the Evolution of Reduced Reactive Aggression in the Context of Human Self-Domestication (2019) · cites this paper as Hublin et al., 2017
In the citing paper
“Some of these facial properties may have been present in the earliest fossils attributed to H. sapiens, like the Jebel Irhoud fossils4, deviating from craniofacial features which emerged in earlier forms of Homo145, and may have become established before some brain-related changes discussed here11,146.”
From A catalog of single nucleotide changes distinguishing modern humans from archaic hominins (2019) · cites this paper as [4]
In the citing paper
“Further evidence of the antiquity of a H. sapiens-like facial morphology is provided by the recent study and redating of new and old fossil material from Jebel Irhoud (Morocco) (Fig. 3f), now placed at ~300 Ka 31”
From The evolutionary history of the human face (2019) · cites this paper as [31]
In the citing paper
“mounting genetic and morphological data support the notion that H. sapiens first appeared in Africa [29][30][31]”
From The evolutionary history of the human face (2019) · cites this paper as [31]
In the citing paper
“Irhoud 1, found in Morocco and recently dated to ~300 Ka 31 , is likely an early member of the Homo sapiens lineage”
From The evolutionary history of the human face (2019) · cites this paper as [31]
In the citing paper
“Compared with early Homo specimens at a comparable stage of dental development, El Sidrón J1 is at least 2.7 years older than a ~2 myr Homo habilis specimen, StW 151, (19)(20)(21)(22) but almost identical in age (7.78 years) to a 315 kyr Homo sapiens specimen from Jebel Irhoud, Morocco (23), that shows a prolonged modern humanlike period of dental developmental (24).”
From The growth pattern of Neandertals, reconstructed from a juvenile skeleton from El Sidrón (Spain) (2017) · cites this paper as [23]
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Citations
Cites
Extensions of the procrustes method for the optimal superimposition of landmarks
Rohlf, F. J., & Slice, D. (1990) · Systematic Zoology · doi:10.2307/2992207 · Cited by 4,289
A high-coverage genome sequence from an archaic denisovan individual
Meyer, M., Kircher, M., Gansauge, M.-T., Li, H., Racimo, F., Mallick, S., Schraiber, J. G., Jay, F., Prüfer, K., de Filippo, C., et al. (2012) · Science · doi:10.1126/science.1224344 · Cited by 1,967
Landmark methods for forms without landmarks: morphometrics of group differences in outline shape
Bookstein, F. L. (1997) · Medical Image Analysis · doi:10.1016/s1361-8415(97)85012-8 · Cited by 1,580
Advances in Geometric Morphometrics
Mitterœcker, P., & Gunz, P. (2009) · Evolutionary Biology · doi:10.1007/s11692-009-9055-x · Cited by 1,416
Pleistocene Homo sapiens from Middle Awash, Ethiopia
White, T. D., Asfaw, B., DeGusta, D., Gilbert, H., Richards, G. D., Suwa, G., & Clark Howell, F. (2003) · Nature · doi:10.1038/nature01669 · Cited by 1,032
Stratigraphic placement and age of modern humans from Kibish, Ethiopia
McDougall, I., Brown, F. H., & Fleagle, J. G. (2005) · Nature · doi:10.1038/nature03258 · Cited by 1,023
THE HUMAN CAREER: HUMAN BIOLOGICAL AND CULTURAL ORIGINS
McHenry, H. M., & Klein, R. G. (1991) · Evolution · doi:10.1111/j.1558-5646.1991.tb04425.x · Cited by 885
Semilandmarks: a method for quantifying curves and surfaces
Gunz, P., & Mitterœcker, P. (2013) · Bollettino del CILEA (CILEA) · doi:10.4404/hystrix-24.1-6292 · Cited by 849
(1982) · Journal of Human Evolution · doi:10.1016/s0047-2484(82)80016-x · Cited by 637
The age of the hominin fossils from Jebel Irhoud, Morocco, and the origins of the Middle Stone Age
Richter, D., Grün, R., Joannes-Boyau, R., Steele, T. E., Amani, F., Rué, M., Fernandes, P., Raynal, J.-P., Geraads, D., Ben-Ncer, A., et al. (2017) · Nature · doi:10.1038/nature22335 · Cited by 576
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Flagged first, then by citations
1 of the 74 references couldn’t be matched to a known paper.
Research works printed but not linked: 28
- Ennouchi, E. Le deuxième crâne de l’homme d’Irhoud. Annales de Paléontologie (Vértébrés) LIV, 117–128 (1968)
- Tixier, J ., Brugal, J.-P ., Tillier, A.-M ., Bruzeket, J . & Hublin, J.-J. in Actes des 1ères Journées Nationales d’Archéologie et du Patrimoine. Préhistoire Vol. 1 149–153 (Société Marocaine d’Archéologie et du Patrimoine, 2001)
- Amani, F. & Geraads, D. Le gisement moustérien du Djebel Irhoud, Maroc: précisions sur la faune et la biochronologie, et description d’un nouveau reste humain. Comptes Rendus à l'Académie des Sciences de Paris 316, 847–852 (1993)
- Ennouchi, E. Un Néanderthalien: l’homme du Jebel Irhoud (Maroc). Anthropologie 66, 279–299 (1962)
- Beucher, S . & Lantuéjoul, C. in International Workshop on Image Processing: Real-Time Edge and Motion Detection 2.1–2.12 (Rennes, France, 1979)
- Sausse, F. La mandibule atlanthropienne de la carrière Thomas I (Casablanca). Anthropologie 79, 81–112 (1975)
- Tillier, A.-M. in Le Squelette moustérien de Kébara 2 97–112 (Centre National de la Recherche Scientifique, Paris, 1991)
- Stewart, T. D. The Neanderthal skeletal remains from Shanidar Cave, Iraq: a summary of findings to date. Proc. Am. Phil. Soc. 121, 121–165 (1977)
- Topinard, P. Les caracteres simiens de la machoire de la Naulette. Rev. Antropol. 15, 385–431 (1886)
- Heim, J. L. Les hommes fossiles de La Ferrassie (Dordogne) et le problem de la definition des Neandertaliens classiques. III. Squelette céphalique. Anthropologie 78, 321–378 (1974)
- de Lumley, M.-A. Les Néandertaliens de la grotte de l’Hortus. Etudes Quaternaires 1, 375–385 (1972)
- Corrain, C. Resti scheletrici umani del ‘Riparo Mezzena’. Memorie del Museo civico di Storia naturale di Verona 16, 97–101 (1968)
- Martin, H. Machoire humaine moustérienne trouvée dans la station de La Quina. L’homme préhistorique 13, 3–21 (1926)
- Vlcˇek, E. Fossile Menschenfunde von Weimar-Ehringsdorf, Weimarer Monographien zur Ur- und Frühgeschichte Vol. 30 (Landesamt für Archäologische Denkmalpflege, 1993)
- Arambourg, C . & Hoffstetter, R. Le gisement de Ternifine Vol. 1. (Masson, 1963)
- Hooton, E.A ., Hencken, H.O & Snow, Ch. E. The ancient Palestinian: Skhu¯ l V reconstruction. 17, 5–10 (American School of Prehistoric Research, 1953)
- Formicola, V. Una mandibola umana dal deposito dell’Epigravettiano finale delle Arene Candite (scavi 197O). Rev. Antropol. 64, 271–278 (1986)
- Odano, A. M. & Riquet, R. Le gisement préhistorique de Dar-es-Soltane 2. Champ de tir de El Menzeh à Rabat (Maroc). Note préliminaire. 2-Étude anthropologique des restes post-atériens. Bull. d’Archéologie Marocaine 11, 25–63 (1978)
- Debénath, A. Nouveaux restes humains atériens du Maroc. CR Acad. Sci. Paris 290, 851–852 (1980)
- Anderson, J. E. In The Prehistory of Nubia Vol. 2 (ed. Wendorf, F. ) 996–1040 (Southern Methodist Univ. Press,1968)
- Crevecoeur, I. From the Nile to the Danube: a comparison of the Nazlet Khater 2 and Oase 1 early modern human mandibles. Anthropologie 42, 203–213 (2004)
- Trinkaus, E. & Svoboda, J. (eds) Early Modern Human Evolution in Central Europe. The People of Dolní Veˇ stonice and Pavlov. The Dolni Vestonice Studies Vol. 12 (Oxford Univ. Press, 2006)
- Sládek, V ., Trinkaus, E ., Hillson, S. W . & Holliday, T. W. The People of the Pavlovian. Skeletal Catalogue and Osteometrics of the Gravettian Fossil Hominids from Dolni Vestonice and Pavlov. The Dolni Vestonice Studies Vol. 5 1–244 (BRNO, 2000)
- Drozdová, E. The evaluation of a rediscovered fragment of human lower jaw, No 21 from Prˇedmostí u Prˇerova. Archeologické rozhledy 53, 452–460 (2001)
- Gambier, D. Vestiges humains du gisement d’Isturitz (Pyrénées—Atlantiques): étude anthropologique et analyse des traces d’action humaine intentionelle. Antiquités nationales 22–23, 9–26 (1990)
- Grine, F. E. & Franzen, J. L. Fossil hominid teeth from the Sangiran Dome (Java, Indonesia). Courier Forschungsinstitut Senckenberg 171, 75–103 (1994)
- Le Cabec, A., Gunz, P., Kupczik, K., Braga, J. & Hublin, J.-J. Anterior tooth root morphology and size in Neanderthals: taxonomic and functional implications. J. Hum. Evol. 64, 169–193 (2013) · DOI
- Turner, C. G. II, Nichol, C. R. & Scott, G. R. Advances in Dental Anthropology (eds Kelley, M., & Larsen, C. ) 13–31 (Wiley Liss, 1991)
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Helm, C., Benoît, J., Mayor, A., Cawthra, H. C., Penn-Clarke, C. R., & Rust, R. (2019) · Proceedings of the Geologists Association · doi:10.1016/j.pgeola.2019.01.001 · Cited by 26
Wu, X., Pei, S., Cai, Y., Tong, H., Xing, S., Jashashvili, T., Carlson, K. J., & Liu, W. (2021) · Journal of Human Evolution · doi:10.1016/j.jhevol.2021.103052 · Cited by 21
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Braucher, R., Oslisly, R., Mesfin, I., Ntoutoume, P. P., & Aster Team (2022) · Philosophical Transactions of the Royal Society B Biological Sciences · doi:10.1098/rstb.2020.0482 · Cited by 20
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Metadata
- Authors
- Hublin, J.-J., Ben-Ncer, A., Bailey, S. E., Freidline, S. E., Neubauer, S., Skinner, M. M., Bergmann, I., Le Cabec, A., Benazzi, S., Harvati, K., & Gunz, P.
- Affiliations
- Collège de France (FR)
- Institut National des Sciences de l'Archéologie et du Patrimoine (MA)
- New York University (US)
- Max Planck Institute for Evolutionary Anthropology (DE)
- University of Kent (GB)
- University of Bologna (IT)
- Senckenberg Centre for Human Evolution and Palaeoenvironment (DE)
- Corresponding author
- Hublin, Jean-Jacques
- Gunz, Philipp
- ORCID
- Venue
- Nature
- Published
- 2017-06-08
- Volume
- 546
- Issue
- 7657
- Pages
- 289-292
- ISSN
- 0028-0836 (ISSN-L)
- 1476-4687
- OpenAlex
- W2623319323
- Cited by
- 1,347
- References
- 74
- Citations by year
- 2026: 84 · 2025: 127 · 2024: 170 · 2023: 131 · 2022: 126 · 2021: 159 · 2020: 117 · 2019: 191 · 2018: 185 · 2017: 55
- FWCI
- 605.49
- Type
- journal article
- Language
- en
- Update notices
- correction (2018-06-13)
- Open access
- green
- doi.org
- Locations
- Nature (publishedVersion)
- PubMed (publishedVersion)
- Kent Academic Repository (University of Kent) (acceptedVersion)
- Leiden Repository (Leiden University) (submittedVersion)
- Data Archiving and Networked Services (DANS) (submittedVersion)
- HAL (submittedVersion)
- RePEc: Research Papers in Economics (submittedVersion)
- Archivio istituzionale della ricerca (Alma Mater Studiorum Università di Bologna) (submittedVersion, other-oa)
- MPG.PuRe (Max Planck Society) (submittedVersion)
- Funders
- Max-Planck-Institut für Evolutionäre Anthropologie
- Topics
- Social Sciences › Anthropology › Pleistocene-Era Hominins and Archaeology
- Social Sciences › Arts and Humanities › Archeology (arts and humanities) › Forensic Anthropology and Bioarchaeology Studies
- Social Sciences › Cultural Studies › Language and cultural evolution
- Keywords
- human origins, Middle Stone Age, endocranial morphology, neurocranium, mandible, dental morphology, facial morphology, thermoluminescence dating
- MeSH terms
- Africa, Animals, Chronology as Topic, Face, Fossils, Humans, Mandible, Morocco, Phylogeny, Skull, Tooth, Hominidae
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