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Fisher, R. A. (1930) · doi:10.5962/bhl.title.27468 · Cited by 17,617

The genetical evolution of social behaviour. I

Hamilton, W. D. (1964) · Journal of Theoretical Biology · doi:10.1016/0022-5193(64)90038-4 · Cited by 15,725

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Trivers, R. L. (2017) · Routledge eBooks · doi:10.4324/9781315129266-7 · Cited by 10,902

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(2017) · Routledge eBooks · doi:10.4324/9781315129266 · Cited by 7,287

Ecology, Sexual Selection, and the Evolution of Mating Systems

Emlen, S. T., & Oring, L. W. (1977) · Science · doi:10.1126/science.327542 · Cited by 6,230

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Alexander, R. D. (1974) · Annual Review of Ecology and Systematics · doi:10.1146/annurev.es.05.110174.001545 · Cited by 3,862

Geometry for the selfish herd

Hamilton, W. D. (1971) · Journal of Theoretical Biology · doi:10.1016/0022-5193(71)90189-5 · Cited by 3,506

Living in Groups

Krause, J., & Ruxton, G. D. (2002) · doi:10.1093/oso/9780198508175.001.0001 · Cited by 3,195

The Social Organisation of Antelope in Relation To Their Ecology

Jarman, P. J. (1974) · Behaviour · doi:10.1163/156853974x00345 · Cited by 2,194

The Jackknife and Bootstrap

Shao, J., & Tu, D. (1995) · Springer Series in Statistics · doi:10.1007/978-1-4612-0795-5 · Cited by 1,985

Local attitudes and elephant spatial distribution in the Bénoué region, Cameroon: implications for human-elephant conflict and conservation

Granados, A. (2011) · Spectrum Research Repository (Concordia University) · Cited by 1

Sexual segregation in spider monkeys in Belize

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Karle, K. A. (2016) · University of North Florida Digital Commons (University of North Florida) · Cited by 1

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Disease control

Walsh, P. D. (2013) · doi:10.1002/9781118520178.ch24 · Cited by 1

Long-term associations and insights on social structure of the Humpback whales in Prince William Sound, AlaskaPreprint

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Macdonald, S. (2014) · doi:10.53846/goediss-4599 · Cited by 1

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Peer Review #2 of "The influence of fish farm activity on the social structure of the common bottlenose dolphin in Sardinia (Italy) (v0.1)"

(2021) · PeerJ · doi:10.7287/peerj.10960v0.1/reviews/2

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(2014) · PeerJ · doi:10.7287/peerj.471v0.1/reviews/1

Peer Review #2 of "Is painting by elephants in zoos as enriching as we are led to believe? (v0.1)"

(2014) · PeerJ · doi:10.7287/peerj.471v0.1/reviews/2

References

(2015) · Elsevier eBooks · doi:10.1016/b978-0-12-802217-7.09991-9

Dinâmica espaço-temporal intragrupo de macacos-prego (Sapajus nigritus) na Mata Atlântica

Luccas, V. R. (2022) · doi:10.11606/t.47.2022.tde-04072022-173010

The Evolutionary Basis for Human Tolerance – Physiological Responses

Spikins, P. (2022) · White Rose University Press eBooks · doi:10.22599/hiddendepths.e

Age structure and grouping behaviour of elephants in Serengeti and Mikumi National Parks in Tanzania.Preprint

Mramba, R. P. (2022) · Research Square · doi:10.21203/rs.3.rs-1744068/v1

Socioconnectomics: connectomics should be enlarged to animal societies to better understand evolutionary processesPreprint

Sueur, C. (2022) · doi:10.31219/osf.io/4ygd7

Realistic modelling of complex systems of biological agents: epidemiology of HIV on complex sexual networks and collective motion of hierarchical herds

Ferdinandy, B. (2018) · Eötvös Loránd Tudományegyetem · doi:10.15476/elte.2017.060

Collective movements during visits to water bodies in wild Asian elephants

Mizuno, K., Ranjeewa, A. D. G., Kutsukake, N., & Padmalal, K. U. K. G. (2023) · Journal of Ethology · doi:10.1007/s10164-023-00790-5

Resting pattern and preferences when lying down at night in a semi-captive population of African elephants

Bonato, M., Padfield, C., Prangnell, B., & Young, D. (2023) · Applied Animal Behaviour Science · doi:10.1016/j.applanim.2023.106052

Social structure and ecological correlates of Indian Blackbuck Antilope cervicapra (Linnaeus, 1758) (Mammalia: Artiodactyla: Bovidae) sociality at Point Calimere Wildlife Sanctuary, India

Arandhara, S., Sathishkumar, S., Gupta, S., & Baskaran, N. (2023) · Journal of Threatened Taxa · doi:10.11609/jott.8451.15.11.24151-24168

Elephant Reproduction

Brown, J. L. (2024) · Elsevier eBooks · doi:10.1016/b978-0-443-21477-6.00115-2

Elephant Vocal Communication

Pardo, M. A. (2024) · Elsevier eBooks · doi:10.1016/b978-0-323-95504-1.00188-5

Analysis of sparse animal social networksPreprint

Mylne, H. K., Abell, J., Beale, C. M., Brent, L. J. N., Bro‐Jørgensen, J., Evans, K. E., Hart, J. D. A., Sakala, D., Simpamba, T., Youldon, D., et al. (2024) · bioRxiv (Cold Spring Harbor Laboratory) · doi:10.1101/2024.10.31.621436

Social and spatial drivers of the multitiered structure of zebra finch social networksPreprint

Zhang, Y., Jiang, X., Aplin, L. M., Wang, D., & Farine, D. R. (2024) · bioRxiv (Cold Spring Harbor Laboratory) · doi:10.1101/2024.11.25.625155

Impact of Population Density on Group Size

Blank, D. A. (2025) · Springer eBooks · doi:10.1007/978-3-031-83504-9_12

Group identity without social interactions?

Ramos‐Fernández, G., Aguilar, S. E. S., Pietrangeli, E., del Toro, C. J., Nicolàs-Arfelis, J. M., Boyer, D., Pinacho‐Guendulain, B., Montiel-Castro, A. J., & Aureli, F. (2025) · Behavioral and Brain Sciences · doi:10.1017/s0140525x24001146

How swarm size affects soldier crab swarming behavior

Feliciani, C., Murakami, H., Tomaru, T., Uesugi, Y., Tanida, S., Nishiyama, Y., Jia, X., & Maeda, T. (2025) · Artificial Life and Robotics · doi:10.1007/s10015-025-01037-x

Dynamic membership drives long-term persistence of sparrow winter social communitiesPreprint

Madsen, A. E., Lyon, B. E., Chaine, A. S., & Shizuka, D. (2025) · bioRxiv (Cold Spring Harbor Laboratory) · doi:10.1101/2025.07.25.666472

African elephants address one another with name-like calls − similar to humans

Pardo, M. (2024) · doi:10.64628/aai.wx4vxf4ct

Comment la sociabilité des éléphants les aide à survivre au braconnage

Goldenberg, S. Z. (2016) · doi:10.64628/aak.h4ywvnsrr

Can elephants retain their social bonds in the face of poaching?

Goldenberg, S. Z. (2015) · doi:10.64628/aai.ce5tnyyxv

Female Asian elephant clans have age-based order but weak dominance resolutionPreprint

Nandini, S., Gautam, H., Keerthipriya, P., & Vidya, T. N. C. (2025) · bioRxiv (Cold Spring Harbor Laboratory) · doi:10.1101/2025.08.22.671838 · Cited by 1

Investigating the patterns of tree cover and density in relation to abiotic and biotic factors in Kruger National Park

Riccardi, T., D’Ammando, G., Wilson, L., Attorre, F., Davies, A. B., Farcomeni, A., MacFadyen, S., Strydom, T., Smit, I. P. J., & Malatesta, L. (2025) · Ecosphere · doi:10.1002/ecs2.70421

Facebook for Geese: The Causes and Consequences of Non-random Social Associations in a Group Forager

Silk, M. J. (2014) · Open Research Exeter (University of Exeter)

Conserving moving species under changing landscapes and climates

Loarie, S. R. (2008) · DukeSpace (Duke University)

Additional file 1 of Patterns of helminth infection in Kenyan elephant populations

King'ori, E. M., Obanda, V., Chiyo, P. I., Soriguer, R. C., Morrondo, P., & Angelone, S. (2020) · Figshare · doi:10.6084/m9.figshare.12005040

The Dominance of Geometric Graph Models in Animal Social NetworksPreprint

Appaw, R. C., Silk, M. J., Rushmore, J., VanderWaal, K. L., Charleston, M., & Fountain‐Jones, N. M. (2025) · bioRxiv (Cold Spring Harbor Laboratory) · doi:10.64898/2025.12.16.694755

Decision-Making on Timing and Route Selection: A Game-Theoretic Approach

Wang, C., & Liu, M. (2025) · IEEE Conference on Decision and Control (CDC) · doi:10.1109/cdc57313.2025.11312638

The Emerging Role of Prolactin as a Biomarker for Reproduction, Health, and Welfare in Wildlife Species: Elephants as a Model

Prado, N. A., & Brown, J. L. (2026) · Advances in experimental medicine and biology · doi:10.1007/978-3-031-87707-0_15

To drink or not to drink? : the influence of resource availability on elephant foraging and habitat selection in a semi-arid savanna

Valls‐Fox, H. (2015) · doi:10.70675/6889e809zb8cfz4a10z9e30z2c893da13df5

Social, ecological and genetic structures of bottlenose dolphins, Tursiops truncatus, in the Normano-Breton gulf and in the North-East Atlantic

Louis, M. (2014) · theses.fr (ABES) · doi:10.70675/ff0d352dz6ad1z459ezb45ezff2f59e55d1a · Cited by 3

Effet de la taille de groupe sur le comportement, la démographie et la dynamique sociale du zèbre des plaines en contexte de prédation

Vitet, C. (2020) · theses.fr (ABES) · doi:10.70675/495d83ddz3597z49b8zb66cz150dc69dd928

Get over here: black capuchin monkeys use long-range vocalizations to adjust the distance between individuals

Luccas, V. R., & Izar, P. (2026) · Primates · doi:10.1007/s10329-026-01260-5

Habitat use differs in released and wild-born Przewalski´s horses of the great Gobi B strictly protected area (Mongolia)

Bernátková, A., Arenas‐Castro, S., Ganbaatar, O., Komárková, M., Sillero, N., Šimek, J., & Ceacero, F. (2026) · European Journal of Wildlife Research · doi:10.1007/s10344-026-02110-x

Pointing with Your Nose:

Bates, L. A., Byrne, R., & Poole, J. (2026) · Cambridge University Press eBooks · doi:10.1017/9781009445696.023

Investigating the adaptive coping mechanisms of rewilded elephants: A comparison of behavioural and physiological variables with wild elephants

Roos, T., Eggeling, T., Garaï, M. E., Boult, V. L., Ganswindt, A., Mitchell, B., & Delsink, A. (2026) · PLoS ONE · doi:10.1371/journal.pone.0348698

Asian elephants (Elephas maximus) demonstrate inhibitory control in a detour task

Hope, S. F., Jacobson, S. L., Dittakul, S., Schmitt, G., PLA-ARD, M., Yindee, M., Angkawanish, T., & Plotnik, J. M. (2026) · PLoS ONE · doi:10.1371/journal.pone.0356549

Paleogenomics and habitat modeling reveal temperate Eurasian origins of woolly rhinoceroses

Xiao-le, L., Ruan, J., Liang, X., Teixeira, R., Zhang, J., Gilardet, A., Zhou, X., Han, Y., Zhao, Z., Yue, H.-Y., et al. (2026) · Science · doi:10.1126/science.aed0430 · Cited by 1

Elephant gestural communication during agonistic interactions: repertoire, intentionality, and goals

Eleuteri, V., Letrari, A., Bates, L. A., Hobaiter, C., & Stoeger, A. S. (2026) · Animal Cognition · doi:10.1007/s10071-026-02108-7

Social context information and acoustic convergence in elephant vocal choruses

Mahon, L., Pardo, M. A., Miron, M., Chemla, E., Granli, P., Njiraini, N., Moss, C. J., Keen, S. C., & Poole, J. H. (2026) · Scientific Reports · doi:10.1038/s41598-026-68525-3

Temporal dynamics of association patterns in Geoffroy's spider monkeys, Ateles geoffroyi: a multiscale approach using temporal multiplex networks

Aguilar, S. E. S., del Toro, C. J., Aureli, F., Pietrangeli, E., & Ramos‐Fernández, G. (2026) · Animal Behaviour · doi:10.1016/j.anbehav.2026.123727

Savannah elephant family dynamics: short- and long-term responses to environmental and social changes

Moss, C. J., Lee, P. C., Njiraini, N., Sayialel, C., & Fishlock, V. (2026) · Animal Behaviour · doi:10.1016/j.anbehav.2026.123733

Reproductive state shapes female social network position in bottlenose dolphins

Methion, S., & López, B. D. (2026) · Animal Behaviour · doi:10.1016/j.anbehav.2026.123745

Behavioral Ecology: Social Organization in Fission–Fusion Societies

Couzin, I. D. (2006) · Current Biology · doi:10.1016/j.cub.2006.02.042 · Cited by 179

A Comparison of Social Organization in Asian Elephants and African Savannah Elephants

de Silva, S., & Wittemyer, G. (2012) · International Journal of Primatology · doi:10.1007/s10764-011-9564-1 · Cited by 134

Where sociality and relatedness diverge: the genetic basis for hierarchical social organization in African elephants

Wittemyer, G., Okello, J. B. A., Rasmussen, H. B., Arctander, P., Nyakaana, S., Douglas‐Hamilton, I., & SIEGISMUND, H. R. (2009) · Proceedings of the Royal Society B Biological Sciences · doi:10.1098/rspb.2009.0941 · Cited by 95

Group size differences may mask underlying similarities in social structure: a comparison of female elephant societies

Nandini, S., Keerthipriya, P., & Vidya, T. N. C. (2017) · Behavioral Ecology · doi:10.1093/beheco/arx135 · Cited by 40

The dynamics of social networks among female Asian elephants

de Silva, S., Ranjeewa, A. D., & Kryazhimskiy, S. (2011) · BMC Ecology · doi:10.1186/1472-6785-11-17 · Cited by 159