Exploring the relationship between personality and social interactions in zoo-housed elephants: incorporation of keeper expertise

From mice to men: What can we learn about personality from animal research?

Gosling, S. D. (2001) · Psychological Bulletin · doi:10.1037/0033-2909.127.1.45 · Cited by 1,683 · see citing works

Updating Animal Welfare Thinking: Moving beyond the “Five Freedoms” towards “A Life Worth Living”

Mellor, D. (2016) · Animals · doi:10.3390/ani6030021 · Cited by 948 · see citing works

Personality in nonhuman primates: a review and evaluation of past research

Freeman, H. D., & Gosling, S. D. (2010) · American Journal of Primatology · doi:10.1002/ajp.20833 · Cited by 356 · see citing works

Allomothering among African elephants

Lee, P. C. (1987) · Animal Behaviour · doi:10.1016/s0003-3472(87)80234-8 · Cited by 253 · see citing works

Observer ratings: Validity and value as a tool for animal welfare research

Meagher, R. K. (2009) · Applied Animal Behaviour Science · doi:10.1016/j.applanim.2009.02.026 · Cited by 247 · see citing works

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Massen, J. J. M., & Koski, S. E. (2013) · Evolution and Human Behavior · doi:10.1016/j.evolhumbehav.2013.08.008 · Cited by 224 · see citing works

Social dominance, seasonal movements, and spatial segregation in African elephants: a contribution to conservation behavior

Wittemyer, G., Getz, W. M., Vollrath, F. W., & Douglas‐Hamilton, I. (2007) · Behavioral Ecology and Sociobiology · doi:10.1007/s00265-007-0432-0 · Cited by 203 · see citing works

Effects of environmental enrichment on reproduction

Carlstead, K., & Shepherdson, D. (1994) · Zoo Biology · doi:10.1002/zoo.1430130507 · Cited by 202 · see citing works

Untitled work

(2009) · Cited by 175 · see citing works

Behavioural profiles are shaped by social experience: when, how and why

Sachser, N., Kaiser, S., & Hennessy, M. B. (2013) · Philosophical Transactions of the Royal Society B Biological Sciences · doi:10.1098/rstb.2012.0344 · Cited by 155 · see citing works

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Positive animal welfare states and encouraging environment-focused and animal-to-animal interactive behaviours

Mellor, D. J. (2014) · New Zealand Veterinary Journal · doi:10.1080/00480169.2014.926800 · Cited by 125 · see citing works

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Archie, E. A., & Chiyo, P. I. (2011) · Molecular Ecology · doi:10.1111/j.1365-294x.2011.05237.x · Cited by 101 · see citing works

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Tetley, C., & O’Hara, S. (2012) · Animal Welfare · doi:10.7120/09627286.21.4.463 · Cited by 99 · see citing works

Personality in Nonhuman Animals

(2017) · Springer eBooks · doi:10.1007/978-3-319-59300-5 · Cited by 90 · see citing works

Early development influences ontogeny of personality types in young laboratory rats

Rödel, H. G., & Meyer, S. (2011) · Developmental Psychobiology · doi:10.1002/dev.20522 · Cited by 73 · see citing works

Applications of Personality to the Management and Conservation of Nonhuman Animals

Powell, D. M., & Gartner, M. C. (2011) · Primatology Monographs · doi:10.1007/978-4-431-53892-9_8 · Cited by 69 · see citing works

Wild female African elephants (Loxodonta africana) exhibit personality traits of leadership and social integration.

Lee, P. C., & Moss, C. J. (2012) · Journal of comparative psychology · doi:10.1037/a0026566 · Cited by 63 · see citing works

Testosterone secretion, musth behaviour and social dominance in captive male Asian elephants living near the equator

Lincoln, G. A., & Ratnasooriya, W. D. (1996) · Reproduction · doi:10.1530/jrf.0.1080107 · Cited by 61 · see citing works

Nocturnal behavior in a group of female African elephants

Wilson, M. L., Bashaw, M. J., Fountain, K., Kieschnick, S., & Maple, T. L. (2006) · Zoo Biology · doi:10.1002/zoo.20084 · Cited by 61 · see citing works

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Grand, A. P., Kuhar, C. W., Leighty, K. A., Bettinger, T. L., & Laudenslager, M. L. (2012) · Applied Animal Behaviour Science · doi:10.1016/j.applanim.2012.09.002 · Cited by 58 · see citing works

Behavior of female African elephants (Loxodonta africana) in captivity

Adams, J., & Berg, J. K. (1980) · Applied Animal Ethology · doi:10.1016/0304-3762(80)90027-9 · Cited by 52 · see citing works

Personality traits of pair members predict pair compatibility and reproductive success in a socially monogamous parrot breeding in captivity

Fox, R. A., & Millam, J. R. (2014) · Zoo Biology · doi:10.1002/zoo.21121 · Cited by 50 · see citing works

Personality assessment in African elephants (Loxodonta africana): Comparing the temporal stability of ethological coding versus trait rating

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Diurnal and nocturnal activity budgets of zoo elephants in an outdoor facility

Horback, K., Miller, L. J., Andrews, J. R. M., & Kuczaj, S. A. (2014) · Zoo Biology · doi:10.1002/zoo.21160 · Cited by 36 · see citing works

Zoo animals: husbandry, welfare and public interactions.

Berger, M., & Corbett, S. (2018) · Cited by 33 · see citing works

Special Relationships between Female Asian Elephants (<i>Elephas maximus</i>) in Zoological Gardens

Garaï, M. E. (1992) · Ethology · doi:10.1111/j.1439-0310.1992.tb00832.x · Cited by 33 · see citing works

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Gartner, M. C., & Weiss, A. (2018) · International Zoo Yearbook · doi:10.1111/izy.12187 · Cited by 32 · see citing works

Personality Assessment and Its Association With Genetic Factors in Captive A sian and A frican Elephants

Yasui, S., Konno, A., Tanaka, M., Idani, G., Ludwig, A., Lieckfeldt, D., & Inoue‐Murayama, M. (2012) · Zoo Biology · doi:10.1002/zoo.21045 · Cited by 31 · see citing works

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Williams, E., Bremner‐Harrison, S., Harvey, N. D., Evison, E., & Yon, L. (2015) · Zoo Biology · doi:10.1002/zoo.21235 · Cited by 26 · see citing works

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Chadwick, C. L., Williams, E., Asher, L., & Yon, L. (2017) · Animal Welfare · doi:10.7120/09627286.26.4.461 · Cited by 24 · see citing works

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Makecha, R., Fad, O., & Kuczaj II, S. A. (2012) · International Journal of Comparative Psychology · doi:10.46867/ijcp.2012.25.01.01 · Cited by 21 · see citing works

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Watters, J. V., Bremner‐Harrison, S., & Powell, D. (2017) · Springer eBooks · doi:10.1007/978-3-319-59300-5_14 · Cited by 10 · see citing works

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Establishing a Link Between Personality and Social Rank in a Group of Bottlenose Dolphins ( Tursiops truncatus )

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Personality and social dynamics of zoo-housed western lowland gorillas (Gorilla gorilla gorilla)

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Untitled work

Wielebnowski (1999) · doi:10.1002/(sici)1098-2361(1999)18:4

Untitled work

Carlstead (1999) · doi:10.1002/(sici)1098-2361(1999)18:1

A systematic review of animal personality in conservation science

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Temperament in Domestic Cats: A Review of Proximate Mechanisms, Methods of Assessment, Its Effects on Human—Cat Relationships, and One Welfare

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Social interactions in zoo-housed elephants: factors affecting social relationships

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Social Dynamics of a Newly Integrated Bachelor Group of Asian Elephants (<i>Elephas maximus</i>): Welfare implications

Schreier, A. L., Readyhough, T. S., Moresco, A., Davis, M., & Joseph, S. (2023) · Journal of Applied Animal Welfare Science · doi:10.1080/10888705.2021.1908141 · Cited by 15 · see citing works

Continuing challenges of elephant captivity: the captive environment, health issues, and welfare implications

Doyle, C., Rally, H., O’Brien, L., Tennison, M., Marino, L., & Jacobs, B. (2024) · PeerJ · doi:10.7717/peerj.18161 · Cited by 14 · see citing works

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Maher, C. J., Gibson, A., Dixon, L. M., & Bacon, H. (2021) · Animals · doi:10.3390/ani11113090 · Cited by 13 · see citing works

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Esattore, B., Šlipogor, V., Saggiomo, L., & Seltmann, M. W. (2021) · Behavioural Processes · doi:10.1016/j.beproc.2021.104361 · Cited by 8 · see citing works

The Integrated Effect of Environmental Conditions and Human Presence on the Behaviour of a Pair of Zoo-Housed Asian Small-Clawed Otters

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Consistency and flexibility of character in free-ranging male African elephants across time, age, and social contexts

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de Sales, A. R., Anastácio, R. S. S., & Pereira, M. J. (2020) · Natural Resources · doi:10.4236/nr.2020.118019 · Cited by 5 · see citing works

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Esattore, B., Villagrán, M., Pluháček, J., Komárková, M., Dušek, A., Kotrba, R., Bartošová, J., & Bartoš, L. (2020) · General and Comparative Endocrinology · doi:10.1016/j.ygcen.2020.113552 · Cited by 4 · see citing works

Personality in Asian elephants (Elephas maximus): Temporal stability and methods of assessment

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The Application Of Animal Personality In Conservation Biology: Investigating A New Tool In North Island Robin (Petroica longipes)

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Dynamics of male African elephant character durability across time and social contextsPreprint

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Personality, Stereotypy, and Responses to Crate Entry in Captive Giant Pandas

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Social Behaviour in Zoo Bachelor Groups: A Case Study of Related South American Fur Seals

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Behaviour and Welfare Impacts of Releasing Elephants from Overnight Tethers: A Zimbabwean Case Study

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Keeper-Dragon Behavioural Differences in Two Komodo Dragon (Varanus komodoensis) Brothers During Training

Pastorino, G. Q., Yadid, Y., Pintus, D., Giovanni, M. D., Cavalleri, A., Preziosi, R. F., & Hashmi, A. (2022) · Acta Scientific Veterinary Sciences · doi:10.31080/asvs.2022.04.0508

Reaktionen von Afrikanischen Elefanten (Loxodonta africana) auf verschiedene Aspekte des Zoomanagements im Hinblick auf Stress und Wohlbefinden unter Berücksichtigung der individuellen Variation

Hambrecht, S. (2020) · doi:10.21248/gups.56647

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Exploring personality assessments of animals in human care: a case study on polar bear ages, hierarchy, and behavior

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Consistency of the ethological coding and rating methods in identifying individual differences in human-managed northwestern gray wolves (Canis lupus occidentalis)

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Blood Oxytocin-like Immunoreactivity, Cortisol, and Caretaker-Rated Personality in Captive Bottlenose Dolphins (Tursiops truncatus): An Exploratory Pilot Study

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Personality assessment in African elephants (Loxodonta africana): Comparing the temporal stability of ethological coding versus trait rating

Horback, K., Miller, L. J., & Kuczaj, S. A. (2013) · Applied Animal Behaviour Science · doi:10.1016/j.applanim.2013.09.009 · Cited by 48 · see citing works

Personality in Asian elephants (Elephas maximus): Temporal stability and methods of assessment

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Social interactions in zoo-housed elephants: factors affecting social relationships

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Personality Assessment in African Elephants (Loxodonta africana)

Horback, K. (2012) · Aquila Digital Community (University of Southern Mississippi)