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Insights from aquaporin structures into drug-resistant sleeping sickness
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Abstract
Trypanosoma brucei is the causal agent of African trypanosomiasis in humans and animals, the latter resulting in significant negative economic impacts in afflicted areas of the world. Resistance has arisen to the trypanocidal drugs pentamidine and melarsoprol through mutations in the aquaglyceroporin TbAQP2 that prevent their uptake. Here, we use cryogenic electron microscopy to determine the structure of TbAQP2 from T. brucei , bound to either the substrate glycerol or to the sleeping sickness drugs, pentamidine or melarsoprol. The drugs bind within the AQP2 channel at a site completely overlapping that of glycerol. Mutations leading to a drug-resistant phenotype were found in the channel lining. Molecular dynamics (MD) simulations showed the channel can be traversed by pentamidine, with a low energy binding site at the centre of the channel, flanked by regions of high energy association at the extracellular and intracellular ends. Drug-resistant TbAQP2 mutants are still predicted to bind pentamidine, but the much weaker binding in the centre of the channel observed in the MD simulations would be insufficient to compensate for the high energy processes of ingress and egress, hence impairing transport at pharmacologically relevant concentrations. The structures of drug-bound TbAQP2 represent a novel paradigm for drug–transporter interactions and are a new mechanism for targeting drugs in pathogens and human cells.
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Jumper, J., Evans, R., Pritzel, A., Green, T., Figurnov, M., Ronneberger, O., Tunyasuvunakool, K., Bates, R., Žídek, A., Potapenko, A., et al. (2021) · Nature · doi:10.1038/s41586-021-03819-2 · Cited by 43,764
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Metadata
- Authors
- Matusevicius, M., Corey, R. A., Gragera, M., Yamashita, K., Sprenger, T., Ungogo, M., Blaza, J. N., Castro-Hartmann, P., Chirgadze, D. Y., Chaitanya Vedithi, S., Afanasyev, P., Melero, R., Warshamanage, R., Gusach, A., Carazo, J.-M., Carrington, M., Blundell, T., Murshudov, G. N., Stansfeld, P. J., Sansom, M. S., De Koning, H. P., Tate, C. G., & Weyand, S.
- Affiliations
- University of Cambridge (GB)
- University of Bristol (GB)
- Centro Nacional de Biotecnología (ES)
- MRC Laboratory of Molecular Biology (GB)
- Ahmadu Bello University (NG)
- University of York (GB)
- Materials and Structure Analysis, Thermofisher Scientific
- Department of Medicine, University of Cambridge, Victor Phillip Dahdaleh Building, Heart and Lung Research Institute
- ETH Zurich (CH)
- Science and Technology Facilities Council (GB)
- University of Warwick (GB)
- University of Oxford (GB)
- University of Glasgow (GB)
- European Bioinformatics Institute (GB)
- Corresponding author
- Weyand, Simone
- ORCID
- Corey, Robin A
- Gragera, Marcos
- Yamashita, Keitaro
- Ungogo, Marzuq
- Blaza, James N
- Chirgadze, Dimitri Y
- Afanasyev, Pavel
- Melero, Roberto
- Warshamanage, Rangana
- Gusach, Anastasiia
- Carazo, José-Maria
- Carrington, Mark
- Blundell, Tom
- Murshudov, Garib N
- Stansfeld, Phillip J
- Sansom, Mark SP
- De Koning, Harry P
- Tate, Christopher G
- Weyand, Simone
- Venue
- eLife
- Publisher
- eLife Sciences Publications, Ltd
- Published
- 2026-01-14
- Volume
- 14
- ISSN
- 2050-084X (ISSN-L)
- PMC
- PMC12803511
- OpenAlex
- W4411933361
- Cited by
- 1
- References
- 1
- Citations by year
- 2026: 1
- Type
- posted content
- Language
- en
- Open access
- gold
- doi.org
- Licence
- Locations
- eLife (publishedVersion, cc-by)
- PubMed (publishedVersion)
- PubMed Central (publishedVersion, cc-by)
- DOAJ (DOAJ: Directory of Open Access Journals) (submittedVersion, cc-by-sa)
- Bristol Research (University of Bristol) (submittedVersion, cc-by)
- Funders
- Wellcome Trust (10.35802/101234, 10.35802/208361, 208398/Z/17/Z, 10.35802/217138)
- Isaac Newton Trust (15.40(a))
- Medical Research Council (U105197215, MRC-DTP PhD fellowship, G0701258)
- Gates Cambridge Trust (Vice-Chancellor's Award)
- Biotechnology and Biological Sciences Research Council (Impact Accelerator Award BB/S506734/1)
- Topics
- Health Sciences › Medicine › Epidemiology › Trypanosoma species research and implications
- Life Sciences › Biochemistry, Genetics and Molecular Biology › Molecular Biology › Ion Transport and Channel Regulation
- Health Sciences › Medicine › Oncology › Drug Transport and Resistance Mechanisms
- Keywords
- Trypanosoma brucei, aquaglyceroporins, sleeping sickness, drug resistance, pentamidine, melarsoprol, glycerol transport, drug uptake, drug-transporter interactions, channel structure, molecular dynamics simulation, cryo-electron microscopy
- MeSH terms
- Binding Sites, Drug Resistance, Glycerol, Humans, Melarsoprol, Mutation, Pentamidine, Protein Binding, Protein Conformation, Trypanocidal Agents, Trypanosoma brucei brucei, Trypanosomiasis, African, Cryoelectron Microscopy, Aquaporin 2, Molecular Dynamics Simulation
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