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Tracing the spread of Celtic languages using ancient genomics
Abstract
Celtic languages are today restricted to Europe’s Atlantic fringes but were widely spoken across the continent during the Iron Age 1–3 . How Celtic achieved this early distribution remains contested 4 , with alternative archaeological and linguistic models proposing different homelands and dispersal routes. Here we analyse genome-wide data from 4,587 ancient individuals, focusing on Bronze and Iron Age Europe, to test the extent to which these models fit reconstructed demographic histories. We identify three main processes shaping European population structure that are relevant to the Celtic dispersal. First, the arrival of Steppe-related ancestry with Bell Beaker-associated groups after around 4400 bp (calibrated years before present or 1950 ce ) across the region in which Celtic and related languages were later spoken. Second, a movement from southwestern Europe into Britain during the Bronze Age (3800–3200 bp ), consistent with scenarios in which Celtic formed along the Atlantic façade. Third, a westward expansion of ancestry related to Late Bronze Age Knovíz–Urnfield-associated populations from eastern central Europe (3200–2800 bp ), affecting France, the British Isles, northeastern Iberia and Italy and later associated with Iron Age Hallstatt and La Tène contexts 5–8 . Although all three movements have parallels with proposed models, we argue that the final movement, from eastern central Europe, is most parsimonious for the spread of Celtic. Although human history does not necessarily follow a parsimonious trajectory 9 and language shifts need not track genetic change, these results delimit the demographic scenarios that are plausible for the emergence of Celtic in prehistoric Europe.
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Cites
Inference of Population Structure using Dense Haplotype Data
Lawson, D. J., Hellenthal, G., Myers, S., & Falush, D. (2012) · PLoS Genetics · doi:10.1371/journal.pgen.1002453 · Cited by 1,188
The genomic history of the Iberian Peninsula over the past 8000 years
Olalde, I., Mallick, S., Patterson, N., Rohland, N., Villalba-Mouco, V., Silva, M., Dulias, K., Edwards, C. J., Gandini, F., Pala, M., et al. (2019) · Science · doi:10.1126/science.aav4040 · Cited by 552
Research works printed but not linked: 53
- Jørgensen, A. R. Celtic. In The Indo-European Language Family: A Phylogenetic Perspective (ed. Olander, T.) 135–151 (Cambridge Univ. Press, 2023). · DOI
- Karl, R. & Stifter, D. The Celtic World: Critical Concepts in Historical Studies (Routledge, 2007).
- Sims-Williams, P. The location of the Celts according to Hecataeus, Herodotus, and other Greek writers. Études Celt. 42, 7–32 (2016).
- Chadwick, N. K. The Celts (Penguin, 1970).
- Schmidt, K. H. The Celts and the ethnogenesis of the Germanic people. Hist. Sprachforsch. 104, 129–152 (1991).
- Haber, M., Mezzavilla, M., Xue, Y. & Tyler-Smith, C. Ancient DNA and the rewriting of human history: be sparing with Occam’s razor. Genome Biol. 17, 1 (2016). · DOI · Paper page
- Stolarek, I. et al. Genetic history of east-central Europe in the first millennium ce. Genome Biol. 24, 173 (2023). · DOI
- Speidel, L. et al. High-resolution genomic history of early medieval Europe. Nature 637, 118–126 (2025). · DOI
- Gretzinger, J. et al. The Anglo-Saxon migration and the formation of the early English gene pool. Nature 610, 112–119 (2022). · DOI
- Antonio, M. L. et al. Stable population structure in Europe since the Iron Age, despite high mobility. eLife 13, e79714 (2024). · DOI
- MacAulay, D. The Celtic Languages (Cambridge Univ. Press, 1992). · DOI
- Ball, M. J. & Muller, N. The Celtic Languages (Routledge, 2012). · DOI
- Haak, W. et al. Massive migration from the steppe was a source for Indo-European languages in Europe. Nature 522, 207–211 (2015). · DOI · Paper page
- Allentoft, M. E. et al. Population genomics of Bronze Age Eurasia. Nature 522, 167–172 (2015). · DOI · Paper page
- Heggarty, P. et al. Language trees with sampled ancestors support a hybrid model for the origin of Indo-European languages. Science 381, eabg0818 (2023). · DOI
- Mallory, J. P. From the steppe to Ireland: the impact of aDNA research. In The Indo-European Puzzle Revisited (eds Kristiansen, K. et al.) 129–145 (Cambridge Univ. Press, 2023). · DOI
- Gallay, A. Bell Beakers today. Pottery, people, culture, symbols in prehistoric Europe. In Proc. International Colloquium Riva del Garda 1998 (ed. Nicolis, F.) 41–57 (Provincia Autonoma di Trento, 2001).
- Cunliffe, B. Setting the scene. In Exploring Celtic Origins (eds Cunliffe, B. & Koch, J. T.) 1–17 (Oxbow Books, 2018). · DOI
- Cunliffe, B. W. Facing the Ocean: The Atlantic and its Peoples (Oxford Univ. Press, 2001).
- Sims-Williams, P. An alternative to ‘Celtic from the East’ and ‘Celtic from the West’. Camb. Archaeol. J. 30, 511–529 (2020). · DOI
- Schrijver, P. Sound change, the Italo-Celtic linguistic unity, and the Italian homeland of Celtic. In Celtic from the West 3 (eds Koch, J. T. & Cunliffe, B.) 489–502 (Oxbow, 2016). · DOI
- Pope, R. Re-approaching Celts: origins, society, and social change. J. Archaeol. Res. 30, 1–67 (2022). · DOI
- Childe, V. G. The Bronze Age (Cambridge Univ. Press, 1930).
- de Hoz, J. Lepontic, Celtiberian, Gaulish and the archaeological evidence. Études Celt. 29, 223–240 (1992).
- Kristiansen, K. Europe Before History (Cambridge Univ. Press, 2000).
- McColl, H. et al. Tracing the spread of Germanic languages using ancient genomics.Nature https://doi.org/10.1038/s41586-025-09064-1 (2026). · DOI
- Yediay, F. E. et al. Ancient genomics support deep divergence between eastern and western Mediterranean Indo-European languages. Preprint at bioRxiv https://doi.org/10.1101/2024.12.02.626332 (2024). · DOI
- Patterson, N. et al. Large-scale migration into Britain during the Middle to Late Bronze Age. Nature 601, 588–594 (2022). · DOI
- Gretzinger, J. et al. Evidence for dynastic succession among early Celtic elites in Central Europe. Nat. Hum. Behav. 8, 1467–1480 (2024). · DOI
- Olalde, I. et al. The Beaker phenomenon and the genomic transformation of northwest Europe. Nature 555, 190–196 (2018). · DOI · Paper page
- Heyd, V. The mobility and migration revolution in 3rd millennium bc Europe. In Rethinking Migrations in Late Prehistoric Eurasia (eds Fernández-Götz, M. et al.) 41–62 (British Academy, 2022). · DOI
- Soetaert, K., Van den Meersche, K. & van Oevelen, D. limSolve: Solving Linear Inverse Models. R package version 1.5.6 https://doi.org/10.32614/cran.package.limsolve (2008). · DOI
- Allentoft, M. E. et al. Population genomics of post-glacial western Eurasia. Nature 625, 301–311 (2024). · DOI
- Papac, L. et al. Dynamic changes in genomic and social structures in third millennium bce central Europe. Sci. Adv. 7, eabi6941 (2021). · DOI
- Olalde, I. et al. Lasting Lower Rhine–Meuse forager ancestry shaped Bell Beaker expansion. Nature 652, 938–946 (2026). · DOI
- Patterson, N. What genetics can say about Iron Age and Bronze Age Britain. In Presenting Counterpoints to the Dominant Terrestrial Narrative of European Prehistory (eds Koch, J. T. et al.) 187–190 (Oxbow Books, 2025). · DOI
- Bretos Ezcurra, M. et al. Genomic insights from a final Bronze Age community buried in a collective tumulus in an Urnfield settlement in northeastern Iberia. Commun. Biol. 8, 1299 (2025). · DOI
- Anthony, D. W. The Horse, the Wheel, and Language: How Bronze-Age Riders from the Eurasian Steppes Shaped the Modern World (Princeton Univ. Press, 2007). · DOI
- Campbell, L. Do languages and genes correlate? Lang. Dyn. Chang. 5, 202–226 (2015). · DOI
- Barbieri, C. et al. A global analysis of matches and mismatches between human genetic and linguistic histories. Proc. Natl Acad. Sci. USA 119, e2122084119 (2022). · DOI
- Sørensen, M. L. S. & Rebay-Salisbury, K. A brief history of urns, urnfields, and burial in the Urnfield culture. In Death and the Body in Bronze Age Europe: From Inhumation to Cremation (eds Sørensen, M. L. S. & Rebay-Salisbury, K.) 15–35 (Cambridge Univ. Press, 2023). · DOI
- Fontijn, D. & Fokkens, H. The Emergence of Early Iron Age ‘Chieftains’ Graves’ in the Southern Netherlands: reconsidering transformations in burial and depositional practices. In The Earlier Iron Age in Britain and the Near Continent (eds Haselgrove, C. & Pope, R.) 354–374 (Oxbow Books, 2007). · DOI
- Sims-Williams, P. Celtic. In The Indo-European Languages (eds Kapović, M. et al.) 352–386 (Routledge, 2017).
- Damgaard, P. B. et al. Improving access to endogenous DNA in ancient bones and teeth. Sci. Rep. 5, 11184 (2015). · DOI · Paper page
- Meyer, M. & Kircher, M. Illumina sequencing library preparation for highly multiplexed target capture and sequencing. Cold Spring Harb. Protoc. 2010, db.prot5448 (2010). · DOI
- Gansauge, M.-T., Aximu-Petri, A., Nagel, S. & Meyer, M. Manual and automated preparation of single-stranded DNA libraries for the sequencing of DNA from ancient biological remains and other sources of highly degraded DNA. Nat. Protoc. 15, 2279–2300 (2020). · DOI · Paper page
- Hanghøj, K., Moltke, I., Andersen, P. A., Manica, A. & Korneliussen, T. S. Fast and accurate relatedness estimation from high-throughput sequencing data in the presence of inbreeding. GigaScience 8, giz034 (2019). · DOI · Paper page
- Rubinacci, S., Ribeiro, D. M., Hofmeister, R. J. & Delaneau, O. Efficient phasing and imputation of low-coverage sequencing data using large reference panels. Nat. Genet. 53, 120–126 (2021). · DOI
- The 1000 Genomes Project Consortium. A global reference for human genetic variation. Nature 526, 68–74 (2015). · DOI · Paper page
- Browning, B. L. & Browning, S. R. Detecting identity by descent and estimating genotype error rates in sequence data. Am. J. Hum. Genet. 93, 840–851 (2013). · DOI · Paper page
- Racimo, F. et al. The spatiotemporal spread of human migrations during the European Holocene. Proc. Natl Acad. Sci. USA 117, 8989–9000 (2020). · DOI
- Pebesma, E. & Graeler, B. gstat: spatial and spatio-temporal geostatistical modelling, prediction and simulation. R package version 2.1.2 https://doi.org/10.32614/cran.package.gstat (2003). · DOI
- Maier, R. et al. On the limits of fitting complex models of population history to f-statistics. eLife 12, e85492 https://doi.org/10.7554/eLife.85492 (2023). · DOI
Metadata
- Authors
- McColl, H., Kroonen, G., Pinotti, T., Barrie, W., Koch, J., Ling, J., Demoule, J.-P., Kristiansen, K., Sikora, M., & Willerslev, E.
- Venue
- Nature
- Published
- 2026-11-04
- ISSN
- 0028-0836
- 1476-4687
- Cited by
- 3
- References
- 2
- Language
- en
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