Large Language Models in Cybersecurity: A Survey of Applications, Vulnerabilities, and Defense Techniques
Open accessAbstract
Large Language Models (LLMs) are transforming cybersecurity by enabling intelligent, adaptive, and automated approaches to threat detection, vulnerability assessment, and incident response. With their advanced language understanding and contextual reasoning, LLMs surpass traditional methods in tackling challenges across domains such as the Internet of Things (IoT), blockchain, and hardware security. This survey provides a comprehensive overview of LLM applications in cybersecurity, focusing on two core areas: (1) the integration of LLMs into key cybersecurity domains, and (2) the vulnerabilities of LLMs themselves, along with mitigation strategies. By synthesizing recent advancements and identifying key limitations, this work offers practical insights and strategic recommendations for leveraging LLMs to build secure, scalable, and future-ready cyber defense systems.
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Yao et al. [20] survey LLM applications across eight cybersecurity domains, including a brief subsection on IoT, but treat agents only tangentially.
“Yao et al. [20] survey LLM applications across eight cybersecurity domains, including a brief subsection on IoT, but treat agents only tangentially.”
From Agentic AI and Large Language Models for Autonomous IoT Cybersecurity: A Systematic Survey, Taxonomy, and Research Roadmap (2026) · cites this paper as [20]
Section 4 says this survey covers eight cybersecurity domains, and §4.8 covers IoT security. The paper mentions an IDS-Agent in §4.8 and multiple LLM agents under “Emerging Multimodal and Autonomous Defenses,” without making agents a main survey domain. The attribution to “Yao et al.” is incorrect for the cited Jaffal et al. paper; the substantive description otherwise broadly fits. (LLM, conf=0.86)
Experimental — accuracy not yet measuredAgents inherit and amplify LLM weaknesses-hallucination, prompt sensitivity, contextual drift-and propagate them across longer decision chains.
“Agents inherit and amplify the weaknesses of the underlying LLM-hallucination, prompt sensitivity, and contextual drift-and propagate them across longer decision chains [20,21].”
From Agentic AI and Large Language Models for Autonomous IoT Cybersecurity: A Systematic Survey, Taxonomy, and Research Roadmap (2026) · cites this paper as [20]
Section 5 identifies hallucinations as an LLM risk, and §5.3 describes how prompts can manipulate LLM outputs. Section 4.5.2 also notes contextual limitations. The paper does not establish that agents amplify these weaknesses or propagate them across longer decision chains; no page numbers are available in our copy. (LLM, conf=0.82)
Experimental — accuracy not yet measuredCybersecurity environments impose distinct and high reliability, auditability, and reproducibility requirements because incorrect or unverifiable automatic actions are commonly found.
“This approach is justified by the recent findings showing that cybersecurity environments impose distinct and high reliability, auditability, and reproducibility requirements because incorrect or unverifiable automatic actions are commonly found [9], [10], [11].”
From The Evolution of Agentic AI in Cybersecurity: From Single LLM Reasoners to Multi-Agent Systems and Autonomous Pipelines (2026) · cites this paper as [11]
The Ethical and Privacy Considerations section calls for LLM-driven security decisions to be transparent, traceable, and open to independent scrutiny. Sections 4.2.8 and 4.5.1 also describe false positives and errors in specific automated security tasks. The paper does not establish a reproducibility requirement or say that incorrect or unverifiable automatic actions are commonly found across cybersecurity environments. (LLM, conf=0.85)
Experimental — accuracy not yet measured
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Towards Explainable Network Intrusion Detection using Large Language Models
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How Effective Are They? Exploring Large Language Model Based Fuzz Driver Generation
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Survey of Vulnerabilities in Large Language Models Revealed by Adversarial AttacksPreprint
Shayegani, E., Mamun, M. A. A., Fu, Y., Zaree, P., Dong, Y., & Abu‐Ghazaleh, N. (2023) · arXiv (Cornell University) · doi:10.48550/arxiv.2310.10844 · Cited by 45 · see citing works
Defending Large Language Models Against Jailbreaking Attacks Through Goal Prioritization
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PassGPT: Password Modeling and (Guided) Generation with Large Language Models
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Spam-T5: Benchmarking Large Language Models for Few-Shot Email Spam DetectionPreprint
Labonne, M., & Moran, S. J. (2023) · arXiv.org · doi:10.48550/arxiv.2304.01238 · Cited by 43 · see citing works
BadPrompt: Backdoor Attacks on Continuous Prompts
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Intelligent Threat Detection—AI-Driven Analysis of Honeypot Data to Counter Cyber Threats
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GPThreats-3: Is Automatic Malware Generation a Threat?
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A Survey and Evaluation of Android-Based Malware Evasion Techniques and Detection Frameworks
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Not The End of Story: An Evaluation of ChatGPT-Driven Vulnerability Description Mappings
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PentestGPT: An LLM-empowered Automatic Penetration Testing Tool
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Privacy and Security Challenges in Large Language Models
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Dual use concerns of generative AI and large language models
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LocalIntel: Generating Organizational Threat Intelligence from Global and Local Cyber Knowledge
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An Adaptive End-to-End IoT Security Framework Using Explainable AI and LLMs
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Harnessing the Power of LLM to Support Binary Taint Analysis
Liu, P.-Z., Sun, C.-N., Zheng, Y.-W., et al. (2023) · arXiv.org · doi:10.48550/arxiv.2310.08275 · Cited by 35
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SLaDe: A Portable Small Language Model Decompiler for Optimized Assembly
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LLMCloudHunter: Harnessing LLMs for Automated Extraction of Detection Rules from Cloud-Based CTI
Schwartz, Y., Ben-Shimol, L., Mimran, D., Elovici, Y., & Shabtai, A. (2025) · International World Wide Web Conference (WWW) · doi:10.1145/3696410.3714798 · Cited by 34 · see citing works
A Comprehensive Survey on Generative AI Solutions in IoT Security
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Unleashing the Power of Generative Model in Recovering Variable Names from Stripped Binary
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Scamming the Scammers: Using ChatGPT to Reply Mails for Wasting Time and Resources
Cambiaso, E., & Caviglione, L. (2023) · Italian Conference on Cybersecurity · doi:10.48550/arxiv.2303.13521 · Cited by 33
Face It Yourselves: An LLM-Based Two-Stage Strategy to Localize Configuration Errors via Logs
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Augmented Datasheets for Speech Datasets and Ethical Decision-Making
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Content-filtering AI systems–limitations, challenges and regulatory approaches
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Review of Generative AI Methods in Cybersecurity
Yigit, Y., Buchanan, W. J., Tehrani, M. G., & Maglaras, L. (2024) · arXiv · doi:10.48550/arxiv.2403.08701 · Cited by 28
DialogBench: Evaluating LLMs as Human-like Dialogue Systems
Ou, J., Lu, J., Liu, C., Tang, Y., Zhang, F., Zhang, D., & Gai, K. (2024) · Conference of the North American Chapter of the Association for Computational Linguistics: Human Language Technologies (NAACL-HLT) · doi:10.18653/v1/2024.naacl-long.341 · Cited by 27 · see citing works
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Large Language Models for Cyber Security: A Systematic Literature Review
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DexBERT: Effective, Task-Agnostic and Fine-Grained Representation Learning of Android Bytecode
Sun, T., Allix, K., Kim, K., Zhou, X., Kim, D., Lo, D., Bissyandé, T. F., & Klein, J. (2023) · IEEE Transactions on Software Engineering · doi:10.1109/tse.2023.3310874 · Cited by 26 · see citing works
Maximizing Penetration Testing Success with Effective Reconnaissance Techniques Using ChatGPT
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A Light Bug Triage Framework for Applying Large Pre-trained Language Model
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Large Language Models are Edge-Case Fuzzers: Testing Deep Learning Libraries via FuzzGPTPreprint
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Linguistic Steganalysis in Few-Shot Scenario
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SoK: Prompt Hacking of Large Language Models
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Pre-training Code Representation with Semantic Flow Graph for Effective Bug Localization
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Complying With Data Handling Requirements in Cloud Storage Systems
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The Ethico-Political Universe of ChatGPT
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Fortifying cloud environments against data breaches: A novel AI-driven security framework
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RepresentThemAll: A Universal Learning Representation of Bug Reports
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Fine-mixing: Mitigating Backdoors in Fine-tuned Language Models
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Examining Zero-Shot Vulnerability Repair with Large Language Models
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Security and Privacy Challenges of AIGC in Metaverse: A Comprehensive Survey
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How Hardened is Your Hardware? Guiding ChatGPT to Generate Secure Hardware Resistant to CWEs
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Large Language Models for Cybersecurity: New Opportunities
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The Hitchhiker's Guide to Program Analysis: A Journey with Large Language ModelsPreprint
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Defending Pre-trained Language Models as Few-shot Learners against Backdoor AttacksPreprint
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Automated CVE Analysis for Threat Prioritization and Impact Prediction
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Generator-Retriever-Generator Approach for Open-Domain Question AnsweringPreprint
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LLMs for Cyber Security: New OpportunitiesPreprint
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Neutralizing Backdoors through Information Conflicts for Large Language ModelsPreprint
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Using LLMs to Automate Threat Intelligence Analysis Workflows in Security Operation CentersPreprint
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Forewarned is Forearmed: A Survey on Large Language Model-based Agents in Autonomous CyberattacksPreprint
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LLM Security and Guardrail Defense Techniques in Web ApplicationsPreprint
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Fixing Hardware Security Bugs with Large Language Models
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Flagged first, then by citations
Research works printed but not linked: 8
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- Authors
- Jaffal, N. O., Alkhanafseh, M., & Mohaisen, D.
- Affiliations
- Birzeit University (PS)
- Department of Computer Science, Birzeit University, Birzeit P.O. Box 14, Palestine
- Department of Computer Science, University of Central Florida, Orlando, FL 32816, USA
- Venue
- AI
- Publisher
- MDPI AG
- Published
- 2025-09-05
- Volume
- 6
- Issue
- 9
- Pages
- 216
- ISSN
- 2673-2688 (ISSN-L)
- OpenAlex
- W4414013482
- Cited by
- 69
- References
- 127
- Citations by year
- 2026: 64 · 2025: 5
- FWCI
- 27.68
- Type
- journal article
- Language
- en
- Open access
- gold
- doi.org
- Licence
- Locations
- AI (publishedVersion, cc-by)
- DOAJ (DOAJ: Directory of Open Access Journals) (submittedVersion, cc-by-sa)
- Topics
- Physical Sciences › Computer Science › Artificial Intelligence › Adversarial Robustness in Machine Learning
- Physical Sciences › Computer Science › Computer Networks and Communications › Network Security and Intrusion Detection
- Physical Sciences › Computer Science › Information Systems › Information and Cyber Security
- Keywords
- large language models, cybersecurity applications, LLM vulnerabilities, threat detection, vulnerability assessment, incident response, Internet of Things security, blockchain security, hardware security
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Registry, retraction, Cites and statistics results are computed, not judged, with one exception: a printed reference that no index lists is read once by a model before it counts as a citation error. Claims are extracted by a model and kept only when their quote appears verbatim in the text; each cited source is then checked, with a model’s help, for whether it says what the claim relies on. Those verdicts can be wrong — read the claim’s quote and the cited source before relying on one.
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