Comparison of microalgal biomasses as functional food ingredients: focus on the composition of cell wall related polysaccharides
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The production of EPS gives an organism the ability to form multicellular structures, which increases harvest efficiency due to the formation of cell aggregates in the medium leading to self-flocculation.
“The production of EPS gives an organism the ability to form multicellular structures, which in turn increases the efficiency with which it can harvest due to the formation of cell aggregates in the medium, which leads to self-flocculation [169].”
From Microalgae-based wastewater treatment: Mechanisms, challenges, recent advances, and future prospects (2023) · cites this paper as [169]
The Introduction (page unavailable) describes EPS secretion, and §3.1 reports extracellular polymers in several microalgae. Neither passage addresses whether EPS enables multicellular structures or cell aggregation, self-flocculation, and more efficient harvesting. Searches for those relationships found no supporting passage in our copy. (LLM, conf=0.72)
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Separation and fractionation of exopolysaccharide from Porphyridium cruentum
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Microalgae-based wastewater treatment: Mechanisms, challenges, recent advances, and future prospects
Abdelfattah, A., Ali, S. S., Ramadan, H., El-Aswar, E. I., Eltawab, R., Ho, S.-H., Elsamahy, T., Li, S., El-Sheekh, M. M., Schagerl, M., et al. (2023) · Environmental Science and Ecotechnology · doi:10.1016/j.ese.2022.100205 · Cited by 812 · see citing works
“The production of EPS gives an organism the ability to form multicellular structures, which in turn increases the efficiency with which it can harvest due to the formation of cell aggregates in the medium, which leads to self-flocculation [169].”
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Levasseur, W., Perré, P., & Pozzobon, V. (2020) · Biotechnology Advances · doi:10.1016/j.biotechadv.2020.107545 · Cited by 551 · see citing works
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“These compounds are present in high concentration and diversity in microalgae [161].”
BackgroundThe potential of microalgae and their biopolymers as structuring ingredients in food: A review
Bernaerts, T. M. M., Gheysen, L., Foubert, I., Hendrickx, M. E., & Van Loey, A. M. (2019) · Biotechnology Advances · doi:10.1016/j.biotechadv.2019.107419 · Cited by 271 · see citing works
Self-citationInnovation in precision fermentation for food ingredients
Augustin, M. A., Hartley, C. J., Maloney, G. R., & Tyndall, S. M. (2023) · Critical Reviews in Food Science and Nutrition · doi:10.1080/10408398.2023.2166014 · Cited by 262 · see citing works
“…(e.g., yeasts, bacteria, filamentous fungi or algae) are used as alternatives to conventional food sources produced by traditional agriculture (Bernaerts et al. 2018; Linder 2019; Ochsenreither et al. 2016; Ritala et al. 2017), both as a source of edible biomass and functional ingredients…”
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“Among others Nannochloropsis, Monodus subterraneus, Phaeodactylum tricornutum, Odontella aurita and Pophyridium cruentum are described [71,72].”
BackgroundLópez-Sánchez, A., Silva-Gálvez, A. L., Aguilar-Juárez, Ó., Senés-Guerrero, C., Orozco-Nunnelly, D. A., Carrillo-Nieves, D., & Gradilla-Hernández, M. S. (2022) · Journal of Environmental Management · doi:10.1016/j.jenvman.2022.114612 · Cited by 180 · see citing works
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Microalgae Polysaccharides: An Alternative Source for Food Production and Sustainable Agriculture
Moreira, J. B., Vaz, B. D. S., Cardias, B. B., Cruz, C. G., de Almeida, A. C. A., Costa, J. A. V., & de Morais, M. G. (2022) · Polysaccharides · doi:10.3390/polysaccharides3020027 · Cited by 137 · see citing works
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“The algal polysaccharides possess potent bioactivities because of their unique physicochemical properties, such as high content of fucose, galactose, uronic acid, and sulfate [154].”
BackgroundCanelli, G., Tarnutzer, C., Carpine, R., Neutsch, L., Bolten, C. J., Dionisi, F., & Mathys, A. (2020) · Frontiers in Nutrition · doi:10.3389/fnut.2020.565996 · Cited by 117 · see citing works
“vulgaris), contained 10–11% carbohydrate, which was comparable to previously reported values of 10–17% for this species (1, 9).”
MethodResultIsolation, structures and biological activities of polysaccharides from Chlorella: a review
Yuan, Q., Li, H., Wei, Z., Lv, K., Gao, C., Liu, Y., & Zhao, L. (2020) · International Journal of Biological Macromolecules · doi:10.1016/j.ijbiomac.2020.09.080 · Cited by 104 · see citing works
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Self-citationEffect of cell disruption methods on the extraction of bioactive metabolites from microalgal biomass
Stirk, W. A., Bálint, P., Vambe, M., Lovász, C., Molnár, Z., Staden, J. V., & Ördög, V. (2019) · Journal of Biotechnology · doi:10.1016/j.jbiotec.2019.10.012 · Cited by 101 · see citing works
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“The intake of this microalgae is safe, and its unique nutritional composition makes it a promising sustainable ingredient for the agrifood industry [134,135].”
BackgroundKey Challenges for the Commercial Expansion of Ingredients from Algae into Human Food Products
Hosseinkhani, N., McCauley, J. I., & Ralph, P. J. (2022) · Algal Research · doi:10.1016/j.algal.2022.102696 · Cited by 98 · see citing works
García-Pérez, P., Cassani, L., Garcia‐Oliveira, P., Xiao, J., Simal‐Gándara, J., Prieto, M. A., & Lucini, L. (2022) · Food Chemistry · doi:10.1016/j.foodchem.2022.135295 · Cited by 97 · see citing works
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Pereira, H., Silva, J., Santos, T., Gangadhar, K. N., Raposo, A., Nunes, C., Coimbra, M. A., Gouveia, L., Barreira, L. A., & Varela, J. C. S. (2019) · Molecules · doi:10.3390/molecules24173192 · Cited by 94 · see citing works
“CTP4 was 15 g/100 g, being similar to the values obtained for other marine strains (Table 1) [25].”
ResultBoukid, F., & Castellari, M. (2021) · Foods · doi:10.3390/foods10010173 · Cited by 97 · see citing works
Arora, N., & Philippidis, G. P. (2021) · Scientific Reports · doi:10.1038/s41598-021-86372-2 · Cited by 90 · see citing works
Promising Sources of Plant-Derived Polyunsaturated Fatty Acids: A Narrative Review
Rizzo, G., Baroni, L., & Lombardo, M. (2023) · International Journal of Environmental Research and Public Health · doi:10.3390/ijerph20031683 · Cited by 89 · see citing works
“, Pophyridium cruentum and Phaeodactylum tricornutum [200,201].”
Innovative and Healthier Dairy Products through the Addition of Microalgae: A Review
Hernández, H., Nunes, M. C., Prista, C., & Raymundo, A. (2022) · Foods · doi:10.3390/foods11050755 · Cited by 91 · see citing works
Molecular and rheological characterization of different cell wall fractions of Porphyridium cruentum
Bernaerts, T., Kyomugasho, C., Looveren, N. V., Gheysen, L., Foubert, I., Hendrickx, M., & Loey, A. M. V. (2018) · Carbohydrate Polymers · doi:10.1016/j.carbpol.2018.05.001 · Cited by 82 · see citing works
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Promising biorefinery products from marine macro and microalgal biomass: A review
Narayanan, M. (2023) · Renewable and Sustainable Energy Reviews · doi:10.1016/j.rser.2023.114081 · Cited by 70 · see citing works
Katiyar, R., Banerjee, S., & Arora, A. (2021) · Biofuels Bioproducts and Biorefining · doi:10.1002/bbb.2187 · Cited by 71 · see citing works
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Reserve, structural and extracellular polysaccharides of Chlorella vulgaris: A holistic approach
Ferreira, A. S., Ferreira, S. S., Correia, A., Vilanova, M., Silva, T. H., Coimbra, M. A., & Nunes, C. (2019) · Algal Research · doi:10.1016/j.algal.2019.101757 · Cited by 69 · see citing works
Microalgal cell disruption: Effect on the bioactivity and rheology of wheat bread
Nunes, M. C., Graça, C., Vlaisavljević, S., Tenreiro, A. M., de Sousa, I. M. N., & Raymundo, A. (2019) · Algal Research · doi:10.1016/j.algal.2019.101749 · Cited by 64 · see citing works
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“Both Tetraselmis strains had an unusually high glucuronic acid content compared with the literature data [76].”
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Hu, Y., Xiao, Y., Liao, K., Leng, Y., & Lu, Q. (2021) · Journal of Chemical Technology & Biotechnology · doi:10.1002/jctb.6850 · Cited by 58 · see citing works
Sha, J., Lu, Z., Ye, J., Wang, G., Hu, Q., Chen, Y., & Zhang, X. (2019) · Algal Research · doi:10.1016/j.algal.2019.101612 · Cited by 56 · see citing works
Canelli, G., Neutsch, L., Carpine, R., Tevere, S., Giuffrida, F., Rohfritsch, Z., Dionisi, F., Bolten, C. J., & Mathys, A. (2019) · Algal Research · doi:10.1016/j.algal.2019.101754 · Cited by 54 · see citing works
“vulgaris significantly differs, ranging in values between 5% and 60% [35].”
BackgroundColusse, G. A., Carneiro, J., Duarte, M. E. R., de Carvalho, J. C., & Noseda, M. D. (2021) · Critical Reviews in Biotechnology · doi:10.1080/07388551.2021.1941750 · Cited by 54 · see citing works
“or structural materials in the cell [20].”
Franco‐Morgado, M., Amador‐Espejo, G. G., Pérez-Cortés, M., & Gutierrez-Uribe, J. A. (2023) · Applied Food Research · doi:10.1016/j.afres.2023.100296 · Cited by 55 · see citing works
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Ahmed, J., & Kumar, V. (2022) · Algal Research · doi:10.1016/j.algal.2021.102617 · Cited by 53 · see citing works
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Kumar, L., Anand, R., Shah, M. P., & Bharadvaja, N. (2022) · Journal of Hazardous Materials Advances · doi:10.1016/j.hazadv.2022.100145 · Cited by 50 · see citing works
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Microalgae biomass deconstruction using green solvents: Challenges and future opportunities
Raj, T., Morya, R., Chandrasekhar, K., Kumar, D., Soam, S., Kumar, R., Patel, A. K., & Kim, S. (2022) · Bioresource Technology · doi:10.1016/j.biortech.2022.128429 · Cited by 44 · see citing works
Thomassen, G., Dael, M. V., You, F., & Passel, S. V. (2019) · Green Chemistry · doi:10.1039/c9gc03071a · Cited by 44 · see citing works
Liu, C., Sun, D., Liu, J., Zhu, J., & Liu, W. (2021) · Bioresources and Bioprocessing · doi:10.1186/s40643-021-00405-2 · Cited by 43 · see citing works
“However, the safety of flocculants is a key consideration, because in addition to producing biodiesel, microalgae cells can also be used for the extraction of microalgae polysaccharides or the production of animal feeds (Bernaerts et al. 2018; Markou and Nerantzis 2013).”
BackgroundComposition and rheological properties of microalgae suspensions: Impact of ultrasound processing
Martínez‐Sanz, M., Garrido, A., Mijlkovic, A., Krona, A., Martínez‐Abad, A., Coll-Marqués, J. M., López‐Rubio, A., & López-Sánchez, P. (2020) · Algal Research · doi:10.1016/j.algal.2020.101960 · Cited by 42 · see citing works
Wang, M., Yin, Z., & Zeng, M. (2022) · Food Hydrocolloids · doi:10.1016/j.foodhyd.2022.107743 · Cited by 40 · see citing works
Secretion of sulfated fucans by diatoms may contribute to marine aggregate formation
Huang, G., Vidal‐Melgosa, S., Sichert, A., Becker, S., Yang, F., Niggemann, J., Iversen, M. H., Cao, Y., & Hehemann, J. (2021) · Limnology and Oceanography · doi:10.1002/lno.11917 · Cited by 41 · see citing works
“Most of the studies on microalgal polysaccharides have been focused on their monosaccharide composition (Bernaerts et al. 2018; Gaignard et al. 2019), which provides important information on the building blocks of the glycan pool but no information on linkages or individual polysaccharide…”
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Lena, G. D., Casini, I., Lucarini, M., del Pulgar, J. S., Aguzzi, A., Caproni, R., Gabrielli, P., & Lombardi‐Boccia, G. (2019) · European Food Research and Technology · doi:10.1007/s00217-019-03346-5 · Cited by 39 · see citing works
López-Pacheco, I. Y., Rodas-Zuluaga, L. I., Cuéllar‐Bermúdez, S. P., Hidalgo-Vázquez, E., Molina-Vázquez, A., Araújo, R. G., Martínez-Ruiz, M., Varjani, S., Barceló, ., Iqbal, H. M. N., et al. (2022) · Marine Drugs · doi:10.3390/md20100601 · Cited by 38 · see citing works
“The cell wall in microalgae represents 2% to 10% of the biomass dry weight [99].”
BackgroundLiu, Y., Wang, Y., Cao, L., Huang, Z., Zhou, Y., Fan, R., & Li, C. (2023) · Foods · doi:10.3390/foods12051067 · Cited by 37 · see citing works
“Previous studies have shown that PSP is bioactive and comprises polymeric carbohydrates composed of long chains of monosaccharide units bound together by glycosidic linkages, and that its multiple biological activities are closely related to monosaccharide composition, diverse glycosidic linkages, molecular weight, an…”
BackgroundImpact of growth medium salinity on galactoxylan exopolysaccharides of Porphyridium purpureum
Ferreira, A. S., Mendonça, I., Póvoa, I., Carvalho, H., Correia, A., Vilanova, M., Silva, T. H., Coimbra, M. A., & Nunes, C. (2021) · Algal Research · doi:10.1016/j.algal.2021.102439 · Cited by 38 · see citing works
Villar-Navarro, E., Garrido-Pérez, C., & Perales, J. A. (2021) · Algal Research · doi:10.1016/j.algal.2021.102389 · Cited by 35 · see citing works
Zhang, L., Selão, T. T., Nixon, P. J., & Norling, B. (2019) · Algal Research · doi:10.1016/j.algal.2019.101702 · Cited by 35 · see citing works
“To concentrate HA from cyanobacterial cultures, 3 volumes of ice-cold ethanol were added to the cleared supernatants and the resulting pellets were dried in air and re-suspended in deionised H2O for further use [33].”
MethodVerspreet, J., Soetemans, L., Gargan, C., Hayes, M., & Bastiaens, L. (2021) · Foods · doi:10.3390/foods10071516 · Cited by 34 · see citing works
“purpureum sample [29] but levels might be higher under different growth conditions, as seen for C.”
BackgroundGomes-Dias, J. S., Teixeira, J. A., & Rocha, C. M. R. (2022) · Food and Bioprocess Technology · doi:10.1007/s11947-022-02812-5 · Cited by 34 · see citing works
“This information increases the relevance of microalgae species such as Tetraselmis chuii and Phaeodactylum tricornutum, whose cell-wall related polysaccharides are reported to be constituted by more than 15% uronic acids and Chlorella vulgaris, whose exopolysaccharide is composed by 10% of uronic acid units (Bernaerts…”
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Oleszek, M., & Krzemińska, I. (2021) · Fuel · doi:10.1016/j.fuel.2021.120676 · Cited by 33 · see citing works
Kumar, V., Jaiswal, K. K., Tomar, M. S., Rajput, V. S., Upadhyay, S., Nanda, M., Vlaskin, M. S., Kumar, S., & KURBATOVA, A. I. (2021) · Biomass Conversion and Biorefinery · doi:10.1007/s13399-021-01906-y · Cited by 33 · see citing works
Castro-Ferreira, C., Gomes-Dias, J. S., Ferreira‐Santos, P., Pereira, R. N., Vicente, A. A., & Rocha, C. M. R. (2021) · Food Hydrocolloids · doi:10.1016/j.foodhyd.2021.107276 · Cited by 31 · see citing works
Spirulina‐based composites for 3D ‐printing
Fredricks, J. L., Iyer, H., McDonald, R., Hsu, J., Jimenez, A. M., & Roumeli, E. (2021) · Journal of Polymer Science · doi:10.1002/pol.20210683 · Cited by 30 · see citing works
Ruiz‐Domínguez, M. C., Salinas-Fuentes, F., Medina, E., Rincón, B., Martı́n, M. A., Gutiérrez, M. C., & Mezquita, P. C. (2022) · Marine Drugs · doi:10.3390/md20020127 · Cited by 31 · see citing works
“tricornutum found in previous studies are variable and highly dependent on strains, culturing conditions, and growth stages [54,56].”
BackgroundAdvanced Mass Balance Characterization and Fractionation of Algal Biomass Composition
Van Wychen, S., Rowland, S. M., Lesco, K. C., Shanta, P. V., Dong, T., & Laurens, L. M. L. (2021) · Journal of Applied Phycology · doi:10.1007/s10811-021-02508-x · Cited by 30 · see citing works
“Using common analytical techniques, across a wide range of species, only 70–80% of the biomass is accounted for (Bernaerts et al. 2018).”
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Microalgae as healthy ingredients for functional foods
Pina-Pérez, M. C., Brück, W. M., Brück, T., & Beyrer, M. (2019) · Elsevier eBooks · doi:10.1016/b978-0-12-816453-2.00004-8 · Cited by 28 · see citing works
Bioremediation of Pyropia-processing wastewater coupled with lipid production using Chlorella sp.
Zheng, S., Chen, S., Zou, S., Yan, Y., Gao, G., He, M., Wang, C., Chen, H., & Wang, Q. (2020) · Bioresource Technology · doi:10.1016/j.biortech.2020.124428 · Cited by 28 · see citing works
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Bernaerts, T., Panozzo, A., Verhaegen, K. A. F., Gheysen, L., Foubert, I., Moldenaers, P., Hendrickx, M., & Loey, A. M. V. (2018) · Food & Function · doi:10.1039/c8fo00261d · Cited by 25 · see citing works
Optimization of spray drying for Chlorella vulgaris by using RSM methodology and maltodextrin
Konar, N., Durmaz, Y., Polat, D. G., & Mert, B. (2022) · Journal of Food Processing and Preservation · doi:10.1111/jfpp.16594 · Cited by 24 · see citing works
Biological detoxification of mycotoxins: Emphasizing the role of algae
Yadavalli, R., Valluru, P., Raj, R., Reddy, C. N., & Mishra, B. (2023) · Algal Research · doi:10.1016/j.algal.2023.103039 · Cited by 24 · see citing works
Light spectra as triggers for sorting improved strains of Tisochrysis lutea
Gao, F., Woolschot, S., Cabanelas, I. T. D., Wijffels, R. H., & Barbosa, M. J. (2020) · Bioresource Technology · doi:10.1016/j.biortech.2020.124434 · Cited by 24 · see citing works
Villar-Navarro, E., Garrido-Pérez, C., & Perales, J. A. (2020) · The Science of The Total Environment · doi:10.1016/j.scitotenv.2020.143057 · Cited by 23 · see citing works
Yin, H.-C., Sui, J.-K., Han, T.-L., Liu, T., & Wang, H. (2022) · Frontiers in Marine Science · doi:10.3389/fmars.2021.836370 · Cited by 23 · see citing works
“7 × 106 Da, and EPS is negatively charged due to the presence of glucuronic acid and sulfate groups (Bernaerts et al., 2018).”
BackgroundMansour, E. A., El-Enin, S. A. A., Hamouda, A. S., & Mahmoud, H. M. (2019) · Environmental Nanotechnology Monitoring & Management · doi:10.1016/j.enmm.2019.100271 · Cited by 23 · see citing works
Influence of different processing techniques on microalgal protein extraction
Moreira, C., Ferreira‐Santos, P., Nunes, R., Carvalho, B., Pereira, H., Teixeira, J. A., & Rocha, C. M. R. (2025) · Algal Research · doi:10.1016/j.algal.2025.103958 · Cited by 22 · see citing works
“Previous studies has been reported a similar composition in terms of proteins with C. vulgaris showing a content of 39.4 %, N. oceanica of 35.1 % and T . chui 31.1 % [7].”
InfluentialZanella, L., & Vianello, F. (2023) · Nutraceuticals · doi:10.3390/nutraceuticals3010010 · Cited by 22 · see citing works
“Synonyms: Porphyridium purpureum = Porphyridium cruentum; Microchloropsis salina = Nannochloropsis salina; Auxenochlorella pyrenoidosa = Chlorella pyrenoidosa; Arthrospira platensis = Spirulina platensis [126].”
BackgroundInfluentialDemir-Yilmaz, I., Schiavone, M., Esvan, J., Guiraud, P., & Formosa‐Dague, C. (2022) · Algal Research · doi:10.1016/j.algal.2023.103102 · Cited by 23 · see citing works
Desjardins, S. M., Laamanen, C. A., Basiliko, N., & Scott, J. A. (2020) · Environmental Reviews · doi:10.1139/er-2020-0004 · Cited by 20 · see citing works
“CWPS and EPS are mainly composed of the monosaccharide’s glucose, 390 galactose, xylose, and mannose (Bernaerts et al. 2018) suggesting a wide variety of possible 391 polysaccharides may be present among the extracted carbohydrates.”
BackgroundAlcaraz, R., Hernández‐Contreras, Á., Iglesias, P., & Hernández, M. D. (2020) · Algal Research · doi:10.1016/j.algal.2020.102167 · Cited by 20 · see citing works
New procedure for separation and analysis of the main components of cyanobacterial EPS
Strieth, D., Schwarz, A. M., Stiefelmaier, J., Erdmann, N., Muffler, K., & Ulber, R. (2021) · Journal of Biotechnology · doi:10.1016/j.jbiotec.2021.01.007 · Cited by 20 · see citing works
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Intracellular and extracellular carbohydrates in microalgae
Synytsya, A., Sushytskyi, L., Saloň, I., Babayeva, T., & Čopı́ková, J. (2023) · Elsevier eBooks · doi:10.1016/b978-0-323-99196-4.00017-6 · Cited by 19 · see citing works
Mollo, L., Drigo, F., Moglie, M., & Norici, A. (2023) · Algal Research · doi:10.1016/j.algal.2023.103256 · Cited by 20 · see citing works
“I. galbana is not covered by calcite scales (coccoliths) as other representatives of the haptophyta group [51,52] and the cell is simply encapsulated by plasma membranes making the algae easily disrupted by mechanical and chemical stress [53].”
An, S. M., Cho, K., Kim, E. S., Ki, H., Choi, G., & Kang, N. S. (2023) · Marine Drugs · doi:10.3390/md21110563 · Cited by 20 · see citing works
“Additionally, it has been reported that the substance contains a variety of beneficial components, including fiber, phytosterols, protein, and minerals [23,24].”
He, Y., Ji, L., Yuan, Y., Rui, D., Li, J., Cheng, P., Sun, L., & Fan, J. (2025) · International Journal of Biological Macromolecules · doi:10.1016/j.ijbiomac.2025.140572 · Cited by 20 · see citing works
Lin, J., Yan, H., Zhao, L., Lǐ, Y., Nahidian, B., Zhu, M., Hu, Q., & Han, D. (2021) · Environmental Microbiology · doi:10.1111/1462-2920.15465 · Cited by 19 · see citing works
- Authors
- Bernaerts, T. M. M., Gheysen, L., Kyomugasho, C., Jamsazzadeh Kermani, Z., Vandionant, S., Foubert, I., Hendrickx, M. E., & Van Loey, A. M.
- Affiliations
- KU Leuven (BE)
- Venue
- Algal Research
- Publisher
- Elsevier BV
- Published
- 2018-06
- Volume
- 32
- Pages
- 150-161
- ISSN
- 2211-9264 (ISSN-L)
- OpenAlex
- W2796053937
- Cited by
- 235
- References
- 109
- Citations by year
- 2026: 16 · 2025: 37 · 2024: 24 · 2023: 30 · 2022: 38 · 2021: 38 · 2020: 25 · 2019: 22 · 2018: 5
- FWCI
- 22.55
- Type
- journal article
- Language
- en
- Open access
- green
- lirias.kuleuven.be
- Licence
- Locations
- Algal Research (publishedVersion)
- Lirias (KU Leuven) (submittedVersion, other-oa)
- Funders
- Fonds Wetenschappelijk Onderzoek
- KU Leuven (KP/14/004)
- Topics
- Life Sciences › Agricultural and Biological Sciences › Aquatic Science › Seaweed-derived Bioactive Compounds
- Life Sciences › Agricultural and Biological Sciences › Food Science › Polysaccharides Composition and Applications
- Physical Sciences › Energy › Renewable Energy, Sustainability and the Environment › Algal biology and biofuel production
- Keywords
- microalgae, functional food ingredients, cell wall polysaccharides, heteropolysaccharide, microalgal biomass composition, Spirulina platensis, Chlorella vulgaris, Porphyridium cruentum, extracellular polymeric substances, uronic acid, sulfate groups, storage polysaccharides, monosaccharide composition, bioactive compounds
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