Choeropsis liberiensis (Artiodactyla: Hippopotamidae)
Open accessAbstract
Choeropsis liberiensis (Morton, 1849), commonly called the pygmy hippopotamus or pygmy hippo, is the only extant species in the genus Choeropsis. A solitary, forest-dwelling ungulate endemic to the Upper Guinean forest ecosystem in West Africa, it is classified as “Endangered” by the International Union for Conservation of Nature and Natural Resources and is a Convention on International Trade in Endangered Species of Wild Flora and Fauna (CITES) Appendix II species. The smaller of the two extant hippopotamids, it is relatively uncommon in zoological facilities and private collections worldwide.
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Written from the paper’s full text.
TL;DR
This Mammalian Species account compiles the taxonomy, morphology, physiology, ecology, behavior, genetics, and conservation status of the pygmy hippopotamus (*Choeropsis liberiensis*), the only extant member of its genus and an Endangered Upper Guinean forest endemic. It emphasizes how much of the species' biology rests on captive animals and indirect field evidence, with wild population size and many ecological parameters still unresolved.
Key findings
- *C. liberiensis* is distinguished from the common hippopotamus by smaller size (wild adult weights recorded at 204 kg for a male and 165 and 170 kg for two females; captive average 160–275 kg, shoulder height 70–80 cm versus >1,500 kg and 130–140 cm in *H. amphibius*), dental formula i 2/1, c 1/1, p 4/4, m 3/3 (total 38), and a distribution restricted to Côte d'Ivoire, Guinea, Liberia, and Sierra Leone.
- Wild population numbers are unknown; estimates range from 2,000–3,000 worldwide to possibly 15,000 for Côte d'Ivoire alone, with country-level figures such as ≤80 ± 10 for Sierra Leone (White 1986), roughly 150 currently for Sierra Leone, 50–150 for two Guinean reserves, and a modeled ~2,000 females and <600 males in the Gola transboundary landscape (128 of 525 examined 1-km² cells occupied).
- Reproduction in captivity: estrous cycle 28–40 days (mean 35 days behaviorally; 27–32 days hormonally), gestation 188–210 days (mean ~200), single calf, birth weights 3–8 kg (mean 5.7 kg); a 1919–1975 review of 219 births found 62% calf survival, with most deaths in the first month and higher birth weight positively correlated with survival past 3 months.
- The captive population at end-2016 comprised 153 males, 229 females, and 9 undetermined-sex animals in 144 institutions, derived from 162 founders; the 2015 population (n = 377) from 60 founders had an average inbreeding coefficient of 0.0600 and gene diversity of 0.9695. Polycystic kidney disease prevalence exceeds 37% in captive animals and may be an X-linked dominant trait.
Methods
- Species account (Mammalian Species) synthesizing published literature, studbook data, and field reports; no new primary data collection.
- Sources include field surveys using transect counts of footprints and dung, camera traps, community/hunter questionnaires, radiotelemetry (one female n = 5 and one male home-range study), and zoo-based morphometric, endocrine, anesthetic, and pathology records.
- Taxonomic treatment follows Boisserie (2005) and Groves and Grubb (2011) in recognizing *Choeropsis* (not *Hexaprotodon*) and two subspecies, *C. l. liberiensis* and *C. l. heslopi*.
Limitations
- Stated: detailed wild information is scarce because in situ research is difficult; most data come from zoo specimens, all of the nominate subspecies. Morphometric measures including body mass are unavailable for wild animals except two females weighed by Hentschel (1990). Population assessments rely on indirect evidence (interviews, bushmeat surveys, footprint/dung transects, camera traps) and require home range, habitat preference, and social behavior data that are lacking; the widely cited ≤3,000 figure originates from gray literature. The radiotelemetry home-range data are biased by translocation, captivity, habituation, small sample size, and only 3 months of post-release tracking. Hematology and biochemistry reference ranges came from >10 but <50 animals of unknown health status and do not meet ASVCP standards. Studbook-based genetic diversity measures assume equal founder relatedness and may contain pedigree errors.
- Evident from the text: many anatomical, physiological, and disease findings rest on single animals or very small samples (e.g., one subadult female eye measurements, one anesthetized male, seven ejaculates), and several ecological claims (diet, activity pattern, calving season) derive from secondhand reports or conflicting accounts rather than direct observation.
Model-written summary. Check it against the paper before relying on it.
Integrity signals
Registry
Cites
Statistics
Claims the paper makes
25 of 266 cited sources checked · 13 supported · 0 flagged · accuracy not yet measured
- Premisecitation
Choeropsis liberiensis is the only extant species in the genus Choeropsis.
“Choeropsis liberiensis (Morton, 1849), commonly called the pygmy hippopotamus or pygmy hippo, is the only extant species in the genus Choeropsis.”
Quote in the paperNot a substantial claim1 of 1 sources checked- cites Boisserie 2005 source
- Premisecitation
Retention of intrinsic adductor muscles in the feet functions to prevent splaying of the toes, an adaptation to walking on muddy substrates.
“Retention of intrinsic adductor muscles in the feet functions to prevent splaying of the toes, an adaptation to walking on muddy substrates (Fisher et al. 2007).”
Quote in the paperNot a substantial claim1 of 1 sources checked- cites Fisher et al. 2007 source
- Premisecitation
The rhomboid muscle group can either originate in the neck or from the occipital bone in C. liberiensis.
“For example, the rhomboid muscle group can either originate in the neck (Campbell 1936;Fisher et al. 2007) or from the occipital bone (Macalister 1873).”
Quote in the paperNot a substantial claim1 of 3 sources checked - Premisecitation
The earliest remains that unequivocally resemble extant hippos appeared in the fossil record in the late Miocene, around 8 million years ago.
“The earliest remains that unequivocally resemble extant hippos appeared in the fossil record in the late Miocene, around 8 million years ago (Weston 2000), and later within this period its range also extended into parts of Eurasia (Boisserie 2005).”
Quote in the paperNot a substantial claim1 of 2 sources checked - Premisecitation
Later within the late Miocene, the range of hippopotamids extended into parts of Eurasia.
“The earliest remains that unequivocally resemble extant hippos appeared in the fossil record in the late Miocene, around 8 million years ago (Weston 2000), and later within this period its range also extended into parts of Eurasia (Boisserie 2005).”
Quote in the paperNot a substantial claim1 of 1 sources checked- cites Boisserie 2005 source
- Premisecitation
The principal threat to survival of C. liberiensis is habitat loss and fragmentation secondary to agriculture, hardwood logging, gold mining, and subsistence farming throughout the Upper Guinean forest ecosystem.
“The principal threat to survival is habitat loss and fragmentation secondary to agriculture, hardwood logging, gold mining, and subsistence farming throughout the Upper Guinean forest ecosystem (Robinson 1971;Verschuren 1983;Anstey 1991b;Roth et al. 2004;Norris et al. 2010;Hoppe-Dominik et al. 2011;Lindsell et al. 2011;Mallon et al. 2011;Fauna and Flora International and Forestry Development Auth”
Quote in the paperSubstantial claim1 of 3 sources checked- cites Robinson 1971 source
The pygmy-hippopotamus passage (labelled Abstract; page unavailable) says forest vegetation is declining, its distribution has become more localised, and agriculture, forestry and mining contribute to its shrinking range. It does not identify habitat loss and fragmentation as the principal threat or specify hardwood logging, gold mining, or subsistence farming; it also names hunting and other factors. (LLM, conf=0.86)
Experimental — accuracy not yet measured - cites Verschuren 1983
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Anstey 1991b
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Camera trap photos of wild C. liberiensis repeatedly show the animals as subjectively thinner than their captive counterparts.
“Camera trap photos of wild C. liberiensis repeatedly show the animals as subjectively thinner than their captive counterparts (Hentschel 1990;Collen et al. 2011;Eshuis 2011;Conway 2013).”
Quote in the paperNot a substantial claim1 of 3 sources checked- cites Hentschel 1990
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Collen et al. 2011 source
The survey account (page not provided in our copy) reports seven camera-trap events that photographed wild pygmy hippopotamuses. It does not describe their apparent thinness or compare them with captive animals, so it supports the photographs but not the claimed comparison. (LLM, conf=0.88)
Experimental — accuracy not yet measured - cites Eshuis 2011
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Observations of radio-collared animals and camera trap studies indicate that activity levels tend to peak from the late afternoon until around midnight and again in the early morning hours.
“Observations of radio-collared animals (Bülow 1987;Hentschel 1990), as well as multiple camera trap studies throughout its range, indicate that activity levels tend to peak from the late afternoon until around midnight and again in the early morning hours (Collen et al. 2011;Eshuis 2011;van Heukelum 2011;Conway 2013).”
Quote in the paperNot a substantial claim1 of 3 sources checked- cites Bülow 1987
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Hentschel 1990
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Collen et al. 2011 source
The paper reports a camera-trap survey of pygmy hippopotamuses in Sapo National Park, so it supports identifying Collen et al. as a camera-trap study. The passage reporting seven photographic events does not give their times or describe peaks in activity from late afternoon to midnight or in the early morning (p. 35–37). (LLM, conf=0.82)
Experimental — accuracy not yet measured
- Premisecitation
Fluorescence in situ hybridization, used to determine the primary sex ratio in ejaculates from males (n = 10), revealed an average of 43% of spermatozoa to have a Y chromosome.
“Fluorescence in situ hybridization, used to determine the primary sex ratio in ejaculates from males (n = 10), revealed an average of 43% of spermatozoa to have a Y chromosome (Saragusty et al. 2012).”
Quote in the paperNot a substantial claim1 of 1 sources checked- cites Saragusty et al. 2012 source
- Premisecitation
C. liberiensis inhabits primary and secondary rainforest and swamps.
“C. liberiensis inhabits primary and secondary rainforest and swamps (Bülow 1987;Hentschel 1990), whereas the common hippopotamus frequents riverine savannahs.”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Bülow 1987
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Hentschel 1990
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Morphometric measures, including body mass, are not available for any animals taken from the wild, except for the two adult females weighed by Hentschel (1990) during his fieldwork in Côte d'Ivoire.
“Morphometric measures, including body mass, are not available for any animals taken from the wild, except for the two adult females weighed by Hentschel (1990) during his fieldwork in Côte d'Ivoire.”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Hentschel 1990
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Several textbooks and gray literature sources from zoological institutions in Europe and the United States give body mass as 160-350 kg and body length of adults as 150-170 cm for C. liberiensis.
“however, several textbooks and gray literature sources from zoological institutions in Europe and the United States (Crandall 1964;Boever 1978;Laws 1984;Taylor and Greenwood 1986;Jarofke 1993;Eulenberger 1995;Nowak 1999;Thompson 2002;Miller 2003Miller , 2007;;Miller et al. 2014;Walzer and Stalder 2014) give body mass as 160-350 kg and body length of adults as 150-170 cm.”
Quote in the paperNot a substantial claim0 of 3 sources checked - Premisecitation
The dental formula of C. liberiensis is i 2/1, c 1/1, p 4/4, m 3/3, total 38.
“The dental formula of C. liberiensis is i 2/1, c 1/1, p 4/4, m 3/3, total 38 (Morton 1844).”
Quote in the paperSubstantial claim0 of 1 sources checked- cites Morton 1844
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
The Lower Guinean species, C. heslopi, was formerly reported in southeastern Nigeria, from the Owerri and Warri provinces of the Niger River Delta eastward from the Cross River.
“The Lower Guinean species, C. heslopi, was formerly reported in southeastern Nigeria, from the Owerri and Warri provinces of the Niger River Delta eastward from the Cross River (Ritchie 1930;Heslop 1945;Corbet 1969).”
Quote in the paperNot a substantial claim0 of 3 sources checked - Premisecitation
The only recorded weights for wild C. liberiensis specimens are from an adult male at 204 kg and two adult females at 165 and 170 kg.
“The only recorded weights for wild specimens are from an adult male at 204 kg (Heslop 1944) and two adult females at 165 and 170 kg (Hentschel 1990).”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Heslop 1944
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Hentschel 1990
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Two subspecies of Choeropsis liberiensis are currently recognized: C. l. heslopi and C. l. liberiensis.
“Two subspecies of Choeropsis liberiensis are currently recognized (Lewison 2011): C. l. heslopi Corbet, 1969:388.”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Lewison 2011
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
The common hippopotamus has two pairs of both maxillary and mandibular incisors, i 2/2, c 1/1, p 4/4, m 3/3, total 40.
“In contrast, the common hippopotamus has two pairs of both maxillary and mandibular incisors, i 2/2, c 1/1, p 4/4, m 3/3, total 40 (Laws 1968).”
Quote in the paperSubstantial claim0 of 1 sources checked- cites Laws 1968 source
- Premisecitation
Shortly after the late 20th century assessment, a survey of the Cestos and Senkwehn rivers reported numerous signs of C. liberiensis presence in both watersheds.
“However, shortly thereafter a survey of the Cestos and Senkwehn rivers reported numerous signs of its presence in both watersheds (Robinson and Suter 1999).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Robinson and Suter 1999
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Asymmetry of the ciliary body gives C. liberiensis a more extensive rear field of vision.
“Asymmetry of the ciliary body also gives C. liberiensis a more extensive rear field of vision (Hegner 1967).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Hegner 1967 source
- Premisecitation
In the early 20th century, Hans Schomburgk found C. liberiensis in many other parts of interior Liberia, including along the Duquea River as well as the Mano and Lofa watersheds in western Liberia near the Sierra Leone border.
“In the early 20th century the German hunter and explorer Hans Schomburgk found C. liberiensis in many other parts of interior Liberia, including along the Duquea River as well as the Mano and Lofa watersheds in western Liberia near the Sierra Leone border (Schomburgk 1912(Schomburgk , 1913a(Schomburgk , 1922)).”
Quote in the paperNot a substantial claim0 of 3 sources checked- cites Schomburgk 1912
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Schomburgk 1913a
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- cites Schomburgk 1922
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- Premisecitation
Historical reports of C. liberiensis existence in Ghana were rejected as erroneous accounts of juvenile common hippos.
“Historical reports of its existence in Ghana were rejected as erroneous accounts of juvenile common hippos (Robinson 2013).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Robinson 2013
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
The kidneys of an adult C. liberiensis are about 75% cortex and 23% medulla; the remaining 2% are the branched tubus maximus and the relatively small renal pelvis.
“The kidneys of an adult C. liberiensis are about 75% cortex and 23% medulla; the remaining 2% are the branched tubus maximus and the relatively small renal pelvis (Maluf 1994).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Maluf 1994 source
- Interpretationcitation
The cranial and dental measurements of C. liberiensis (n = 20-30) are consistent with the morphology of browsers.
“The cranial and dental measurements (cm, ranges) of C. liberiensis (n = 20-30) are consistent with the morphology of browsers (Weston 2000(Weston , 2003)):”
Quote in the paperSubstantial claim0 of 2 sources checked - Premisecitation
Recent confirmed reports substantiate the presence of C. liberiensis in the Ziama and Diécké forests and the Mont Béro Reserve.
“Recent confirmed reports substantiate its presence in the Ziama and Diécké forests and the Mont Béro Reserve (Mallon et al. 2011).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Mallon et al. 2011
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Within Côte d'Ivoire, C. liberiensis has not been recorded farther east than the Bandama River and Azagny National Park, about 80 km west of Abidjan.
“Within Côte d'Ivoire C. liberiensis has not been recorded farther east than the Bandama River (Sidney 1965) and Azagny National Park (Bülow 1987), about 80 km west of Abidjan.”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Sidney 1965
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Bülow 1987
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
The cardiovascular anatomy of C. liberiensis is not different from that of other large mammals.
“The cardiovascular anatomy of C. liberiensis is not different from that of other large mammals (Macalister 1873;Macdonald 1988).”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Macalister 1873
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Macdonald 1988 source
- Interpretationcitation
The large number of short nephrons in C. liberiensis is believed to have allowed the evolution of a bigger animal, albeit with a limited ability to produce concentrated urine.
“The large number of short nephrons is believed to have allowed the evolution of a bigger animal, albeit with a limited ability to produce concentrated urine (Maluf 1978(Maluf , 1994)).”
Quote in the paperNot a substantial claim0 of 2 sources checked - Premisenone given
The pylorus connects to a simple linear small intestine.
“The pylorus connects to a simple linear small intestine.”
Quote in the paperNot a substantial claim - Premisecitation
Hippos are called pseudo-ruminants or ruminant-like, with microbial fermentation in the fore stomachs supporting digestion through the production of volatile fatty acids and essential vitamins.
“Hippos are called “pseudo-ruminants” or ruminant-like, with microbial fermentation in the fore stomachs supporting digestion through the production of volatile fatty acids and essential vitamins (Langer 1975;Macdonald and Hartman 1983).”
Quote in the paperNot a substantial claim0 of 2 sources checked - Premisecitation
The blind sacs, ventriculus, and connecting chamber of C. liberiensis are further subdivided by a series of semilunar folds, 1-3 cm in height, derived from the tunica muscularis.
“The blind sacs, ventriculus, and connecting chamber are further subdivided by a series of semilunar folds, 1-3 cm in height, derived from the tunica muscularis (Langer 1975;Macdonald and Hartman 1983).”
Quote in the paperNot a substantial claim0 of 2 sources checked - Interpretationcitation
Southwestern Côte d'Ivoire represents the easternmost range of C. liberiensis.
“Southwestern Côte d'Ivoire represents the easternmost range of C. liberiensis.”
Quote in the paperNot a substantial claim - Premisecitation
The kidney of an adult C. liberiensis is about 133 mm long by 92 mm wide with a thickness of 33 mm.
“The kidney of an adult is about 133 mm long by 92 mm wide with a thickness of 33 mm (Maluf 1994).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Maluf 1994 source
- Premisecitation
The lateral placement of eyes and a tapering head shape facilitate a binocular field of vision when the mouth is gaping open, allowing the animal to advance toward a threat while maintaining visual contact.
“The lateral placement of eyes and a tapering head shape also facilitate a binocular field of vision when the mouth is gaping open, allowing the animal to advance toward a threat while maintaining visual contact (Van den Bergh 1971).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Van den Bergh 1971
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
A distinct border along the lesser curvature marks the separation between the nonglandular connecting chamber and glandular epithelium of the stomach.
“A distinct border along the lesser curvature marks the separation between the nonglandular connecting chamber and glandular epithelium of the stomach (Langer 1975).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Langer 1975 source
- Premisecitation
Hunter and bushmeat market interviews conducted in Nimba and Grand Bassa counties and near Sapo National Park revealed a declining trend in C. liberiensis numbers over the last decade.
“Hunter and bushmeat market interviews conducted in Nimba and Grand Bassa counties and near Sapo National Park revealed a declining trend in C. liberiensis numbers over the last decade (Greengrass 2011).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Greengrass 2011
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
The hair structure of C. liberiensis is similar to that of stiff bristles, comprised of multiple keratin strands that are split at the ends, especially those of lip and tail.
“The hair structure, similar to that of stiff bristles, is comprised of multiple keratin strands that are split at the ends, especially those of lip and tail (Lochte 1951).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Lochte 1951
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisenone given
The esophagus of C. liberiensis opens into the dorsal part of the visceral blind sac.
“The esophagus opens into the dorsal part of the visceral blind sac.”
Quote in the paperNot a substantial claim - Premisecitation
Branching of the tubus maximus and the absence of renculi within a lobed kidney are features found only in hippopotamids, making their renal anatomy unique among the Artiodactyla.
“Branching of the tubus maximus and the absence of renculi within a lobed kidney are features found only in hippopotamids, making their renal anatomy unique among the Artiodactyla (Maluf 1994).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Maluf 1994 source
- Premisecitation
As in ruminants, a fornix-like blind sac in the glandular stomach, lined with nonglandular epithelium, exposes the stomach contents to digestive enzymes.
“As in ruminants, a fornix-like blind sac in the glandular stomach, lined with nonglandular epithelium, exposes the stomach contents to digestive enzymes (Langer 1975).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Langer 1975 source
- Premisecitation
Historically the most westward report of C. liberiensis is from Forécariah, Guinea, about 60 km east of Conakry.
“Historically its most westward report is from Forécariah, Guinea, about 60 km east of Conakry (Dekeyser 1955).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Dekeyser 1955
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisenone given
Insensible water loss from the skin of a nonsedated female C. liberiensis at a temperature range of 29-35°C was between 6.0 and 22.5 mg/5 cm2/10 min, depending on the areas of the body.
“Insensible water loss from the skin of a nonsedated female C. liberiensis at a temperature range of 29-35°C was between 6.0 and 22.5 mg/5 cm 2 /10 min, depending on the areas of the body.”
Quote in the paperNot a substantial claim - Interpretationcitation
Based on a number of field surveys and recent camera trap photos, Taï National Park probably holds many C. liberiensis.
“Based on a number of field surveys and recent camera trap photos, Taï National Park probably holds many C. liberiensis (Roth and Merz 1986;Hentschel 1990;Roth et al. 2004;Eshuis 2011;van Heukelum 2011;Hoppe-Dominik et al. 2011;Bogui et al. 2016).”
Quote in the paperNot a substantial claim0 of 3 sources checked- cites Roth and Merz 1986
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Hentschel 1990
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Bogui et al. 2016
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Carbon dating indicates that the Cypriot population went extinct about 10,500 years ago.
“Carbon dating indicates that the Cypriot population went extinct about 10,500 years ago (Simmons 1999), while the Malagasy population probably survived until about 2,000 years ago (MacPhee and Burney 1991).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Simmons 1999 source
- Premisecitation
Each kidney of C. liberiensis is supplied with cranial and caudal renal arteries that branch into several perforating arteries, which in turn supply their respective lobes.
“Each kidney is supplied with cranial and caudal renal arteries that branch into several perforating arteries, which in turn supply their respective lobes (Maluf 1978;1994).”
Quote in the paperNot a substantial claim0 of 2 sources checked - Premisecitation
The earliest confirmation of C. liberiensis from Liberia is from the St. Paul River and the Duquea River, a tributary of the Junk River.
“The earliest confirmation of this species from Liberia is from the St. Paul River and the Duquea River, a tributary of the Junk River (Büttikofer 1890).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Büttikofer 1890
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Surveys have also confirmed the presence of C. liberiensis in Cavally and Goin Débé Classified Forests.
“Surveys have also confirmed its presence in Cavally and Goin Débé Classified Forests (Mallon et al. 2011).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Mallon et al. 2011
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
The northernmost extent of C. liberiensis in Côte d'Ivoire is the area around Mt. Nimba in the upper Cavally River valley.
“The northernmost extent of C. liberiensis in Côte d'Ivoire is the area around Mt. Nimba in the upper Cavally River valley (Roth et al. 2004).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Roth et al. 2004 source
- Premisecitation
In Guinea, C. liberiensis occurs primarily in the forested regions of the southeast.
“In Guinea, C. liberiensis occurs primarily in the forested regions of the southeast (Sidney 1965).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Sidney 1965
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
In the first one-half of the 20th century, C. liberiensis was widely distributed in the eastern part of Sierra Leone.
“In the first one-half of the 20th century, C. liberiensis was widely distributed in the eastern part of Sierra Leone (Sidney 1965).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Sidney 1965
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Later anatomical studies have revealed several variations in topographical muscle anatomy for C. liberiensis.
“However, later anatomical studies have revealed several variations in topographical muscle anatomy for C. liberiensis.”
Quote in the paperNot a substantial claim - Premisecitation
The myology of C. liberiensis is generally similar to that of the common hippopotamus, as detailed from an approximately 8-week-old female specimen.
“The myology of C. liberiensis is generally similar to that of the common hippopotamus, as detailed from an approximately 8-week-old female specimen (Macalister 1873).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Macalister 1873
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
In the adult C. liberiensis, the papillae increase the functional surface area of the fore stomachs by 3.32 times compared to that of a smooth wall.
“In the adult C. liberiensis, the papillae increase the functional surface area of the fore stomachs by 3.32 times compared to that of a smooth wall (Langer 1975).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Langer 1975 source
- Premisecitation
The presence of C. liberiensis throughout both coastal and interior regions of Liberia was confirmed as recently as the 1960s.
“Its presence throughout both coastal and interior regions of Liberia was confirmed as recently as the 1960s (Sidney 1965).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Sidney 1965
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
C. liberiensis is also known from the Loma Mountains in northeastern Sierra Leone.
“It is also known from the Loma Mountains in northeastern Sierra Leone (Mallon et al. 2011).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Mallon et al. 2011
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Interpretationcitation
If the Nigerian species persists, the only remaining habitat where it could possibly occur is the Upper Orashi Forest Reserve.
“If the Nigerian species persists, the only remaining habitat where it could possibly occur is the Upper Orashi Forest Reserve (Robinson 2013).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Robinson 2013
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premiseown data
In both Tiwai Island and Gola Forest Reserve Complex, presence of C. liberiensis was confirmed via camera trap photos, personal observations, and indirect methods such as tracks and dung.
“In both locations its presence was confirmed via camera trap photos, personal observations, and by indirect methods such as tracks and dung.”
Quote in the paperNot a substantial claim - Interpretationreasoning
Liberia represents the core of the distribution of C. liberiensis.
“Liberia represents the core of its distribution, as reflected in the specific name, C. liberiensis.”
Quote in the paperNot a substantial claim - Premisecitation
The Malagasy population probably survived until about 2,000 years ago.
“Carbon dating indicates that the Cypriot population went extinct about 10,500 years ago (Simmons 1999), while the Malagasy population probably survived until about 2,000 years ago (MacPhee and Burney 1991).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites MacPhee and Burney 1991 source
- Premisecitation
For Guinea, historical surveys estimated the two areas together might hold 50-150 individuals.
“For Guinea, the only historical surveys from Massif du Ziama and Diécké Forest Reserves estimated the two areas together might hold 50-150 individuals (Bützler 1994); there are no current estimates of C. liberiensis numbers from Guinea (Mallon et al. 2011).”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Bützler 1994
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Mallon et al. 2011
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
The habitat of C. liberiensis is centered on areas with plentiful stream banks, swamps, and low-lying areas with year-round standing water.
“The habitat of C. liberiensis (Fig. 4) is centered on areas with plentiful stream banks, swamps, and low-lying areas with year-round standing water (Bigourdan and Prunier 1937).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Bigourdan and Prunier 1937
- Premisecitation
A review of all known births from 1919 to 1975 (n = 219) revealed an overall calf survival rate of 62%.
“A review of all known births from 1919 to 1975 (n = 219) revealed an overall calf survival rate of 62%, with the great majority of deaths occurring within the first month of life (Leutenegger 1978).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Leutenegger 1978 source
- Premisecitation
The great majority of C. liberiensis calf deaths occur within the first month of life.
“A review of all known births from 1919 to 1975 (n = 219) revealed an overall calf survival rate of 62%, with the great majority of deaths occurring within the first month of life (Leutenegger 1978).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Leutenegger 1978 source
- Premisecitation
The interareolar villi have prominent interdigitation between the fetal capillaries and maternal trophoblast, a feature thought to enhance gas exchange across the placenta.
“The latter have prominent interdigitation between the fetal capillaries and maternal trophoblast, a feature thought to enhance gas exchange across the placenta (Macdonald and Bosma 1985;Macdonald 2007).”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Macdonald and Bosma 1985 source
- cites Macdonald 2007
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Investigation of DNA samples from C. liberiensis revealed > 1,600 single-nucleotide polymorphisms (SNPs) suitable for population genetic analysis.
“Investigation of DNA samples from C. liberiensis revealed > 1,600 single-nucleotide polymorphisms (SNPs) suitable for population genetic analysis (Senn et al. 2014).”
Quote in the paperSubstantial claim0 of 1 sources checked- cites Senn et al. 2014 source
- Premisecitation
Median blood gas parameters for three adult C. liberiensis were: pH 7.43; partial pressure of carbon dioxide 56 mmHg; partial pressure of oxygen 40 mmHg; bicarbonate 35.9 mmol/l.
“Median blood gas parameters for three adult C. liberiensis were: pH, 7.43; partial pressure of carbon dioxide (mmHg), 56; partial pressure of oxygen (mmHg), 40; bicarbonate (mmol/l), 35.9 (Bouts et al. 2012).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Bouts et al. 2012 source
- Interpretationreasoning
Estimating local population densities from these indirect methodologies requires knowledge of home range, habitat preference, and social behavior, data that are currently lacking.
“Estimating local population densities from these indirect methodologies requires knowledge of home range, habitat preference, and social behavior, data that are currently lacking.”
Quote in the paperSubstantial claim - Premisecitation
Birth weights of C. liberiensis range from 3 to 8 kg with a mean of 5.7 kg.
“Birth weights range from 3 to 8 kg with a mean of 5.7 kg (Steinmetz 1937;Roth 1962;Nowak 1999;Stroman and Slaughter 1972;Lang 1975;Leutenegger 1978;Greed 1983).”
Quote in the paperNot a substantial claim0 of 3 sources checked- cites Steinmetz 1937
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Roth 1962
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Nowak 1999 source
- Premisecitation
For Sierra Leone, population surveys concluded that the countrywide total was no more than 80 ± 10 animals.
“For Sierra Leone, population surveys concluded that the countrywide total was no more than 80 ± 10 animals, with Tiwai Island near Gola W National Park holding as many as 10 animals (White 1986).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites White 1986
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisenone given
The population data reported in all of these assessments are based on indirect evidence including interviews, bushmeat surveys, transect counts, and camera trap photographs.
“The population data reported in all of these assessments are based on indirect evidence including: interviews with local natives (primarily hunters and farmers) and logging company employees; bushmeat surveys; transect counts of footprints and dung; and camera trap photographs.”
Quote in the paperNot a substantial claim - Premisenone given
Endoparasites have only been reported from captive C. liberiensis.
“Endoparasiteshave only been reported from captive Choeropsis liberiensis.”
Quote in the paperNot a substantial claim - Premisecitation
The first physiological variables for C. liberiensis were reported from an anesthetized female: heart rate 96-106 beats/min, rectal temperature 35.8-38.9°C.
“The first physiological variables for C. liberiensis were reported from an anesthetized female C. liberiensis: heart rate 96-106 beats/min, rectal temperature (°C) 35.8-38.9(Franz et al. 1978).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Franz et al. 1978 source
- Premisecitation
The most recent population estimate for Taï National Park is 2,000.
“The most recent population estimate for Taï National Park is 2,000 (Mallon et al. 2011).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Mallon et al. 2011
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisenone given
Nothing is known about the genetic structure or diversity of the wild C. liberiensis populations.
“Nothing is known about the genetic structure or diversity of the wild C. liberiensis populations.”
Quote in the paperSubstantial claim - Premisecitation
Metabolite levels of pregnanediol in fecal samples steadily increased during pregnancy, reaching concentrations markedly above luteal phase levels in the second one-half of gestation.
“Metabolite levels of preganediol in fecal samples steadily increased during pregnancy, reaching concentrations markedly above luteal phase levels in the second one-half of gestation (Flacke et al. 2017a).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Flacke et al. 2017a
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
C. liberiensis will also eat crop plants, including sweet potato leaves, okra, and cassava, growing in village fields and palm plantations.
“C. liberiensis will also eat crop plants, including sweet potato leaves (Ipomoea batatas-Convolvulaceae), okra (Abelmoschus esculentus-Malvaceae), and cassava (Manihot esculenta-Euphorbiaceae), growing in village fields and palm plantations (Robinson 1970;Bülow 1987;Hentschel 1990).”
Quote in the paperNot a substantial claim0 of 3 sources checked- cites Robinson 1970
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Bülow 1987
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Hentschel 1990
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
C. liberiensis (n = 12) consumed 150 different plant species as food sources in its natural environment in Taï National Park, Côte d'Ivoire.
“Choeropsis liberiensis (n = 12) consumed 150 different plant species as food sources in its natural environment in Taï National Park, Côte d'Ivoire (Hentschel 1990).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Hentschel 1990
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Many young C. liberiensis in captivity have drowned, even in very shallow water.
“Successful parturition in C. liberiensis takes place on land; many young animals in captivity have drowned, even in very shallow water (Stroman and Slaughter 1972;Lang 1975;von Houwald et al. 2007).”
Quote in the paperNot a substantial claim0 of 3 sources checked- cites Stroman and Slaughter 1972 source
- cites Lang 1975
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites von Houwald et al. 2007
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisenone given
C. liberiensis exhibits the unique behavior of backing against vertical surfaces and with a series of rapid, propeller-like tail movements, spreading a mixture of urine and fecal material.
“Choeropsis liberiensis exhibits the unique behavior of backing against vertical surfaces and with a series of rapid, propeller-like tail movements, spreading a mixture of urine and fecal material.”
Quote in the paperNot a substantial claim - Premisecitation
In Côte d'Ivoire, indirect population surveys during 1977-1983 and 1995-2004 indicated a declining trend for C. liberiensis.
“In Côte d'Ivoire, indirect population surveys during 1977-1983 and 1995-2004 indicated a declining trend for C. liberiensis (Hoppe-Dominik et al. 2011).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Hoppe-Dominik et al. 2011 source
- Premisecitation
Respiratory rate for conscious C. liberiensis is reported at 10-16 breaths/min.
“Respiratory rate (breaths/min) for conscious C. liberiensis is reported at 10-16 (Pearce et al. 1985).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Pearce et al. 1985 source
- Premisereasoning
Population numbers for wild C. liberiensis are unknown.
“Population numbers for wild Choeropsis liberiensis are unknown; current assessments are largely unsubstantiated and based on qualitative data.”
Quote in the paperNot a substantial claim - Interpretationreasoning
These circumstances, the small sample size, and the fact that post-release tracking in Azagny was limited to 3 months likely biased the data concerning individual home range size, as well as the overlap of male and female home ranges.
“These circumstances, the small sample size, and the fact that post-release tracking in Azagny was limited to 3 months likely biased the data concerning individual home range size, as well as the overlap of male and female home ranges.”
Quote in the paperSubstantial claim - Premisecitation
Quantitative information on wild C. liberiensis population numbers is mostly lacking.
“Quantitative information is mostly lacking, numbers being very difficult to estimate accurately and often based on informed guesswork or an association of suitable habitat with the possibility of presence (Mallon et al. 2011;Hillers et al. 2016).”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Mallon et al. 2011
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Hillers et al. 2016
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Preferred food plants have higher sodium (Na+) content than other plant species consumed by C. liberiensis.
“Preferred food plants have higher sodium (Na + ) content than other plant species consumed by C. liberiensis (Hentschel 1990).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Hentschel 1990
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Successful parturition in C. liberiensis takes place on land.
“Successful parturition in C. liberiensis takes place on land; many young animals in captivity have drowned, even in very shallow water (Stroman and Slaughter 1972;Lang 1975;von Houwald et al. 2007).”
Quote in the paperNot a substantial claim0 of 3 sources checked- cites Stroman and Slaughter 1972 source
- cites Lang 1975
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites von Houwald et al. 2007
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Estimates of 0.8-2.5 C. liberiensis per km2 of suitable habitat were extrapolated to generate abundance data for Taï National Park plus buffer zones: 12,000 in 1982; 5,000 in 1997.
“Estimates of 0.8-2.5 per km 2 of suitable habitat were extrapolated to generate abundance data for Taï National Park plus buffer zones: 12,000 in 1982; 5,000 in 1997 (Roth et al. 2004).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Roth et al. 2004 source
- Premisecitation
In captivity, C. liberiensis often becomes overweight.
“In captivity, C. liberiensis often becomes overweight (Flacke et al. 2015).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Flacke et al. 2015 source
- Premisecitation
Hippopotamids have comparatively low energy requirements and basal metabolic rates than similarly sized ruminants.
“Hippopotamids have comparatively low energy requirements and basal metabolic rates than similarly sized ruminants (Clauss et al. 2004), so even a small excess in energy intake can lead to weight gain.”
Quote in the paperSubstantial claim0 of 1 sources checked- cites Clauss et al. 2004 source
- Premisecitation
Progesterone levels below 500 pmol/l were noted every 27-32 days, consistent with physiological estrus.
“Progesterone levels below 500 pmol/l were noted every 27-32 days, consistent with physiological estrus (Dathe and Kuckelkorn 1989).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Dathe and Kuckelkorn 1989
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Accumulations of yellow-brown pigment occur in some trophoblasts, including in some of the villous tips, a histologic feature distinctly different from other placentas in the Artiodactyla.
“Accumulations of yellowbrown pigment occur in some trophoblasts, including in some of the villous tips, a histologic feature distinctly different from other placentas in the Artiodactyla (Benirschke 2007).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Benirschke 2007
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Metabolite levels of progesterone, as measured in the skin secretions of two females for six consecutive months, were 104-2,662 pmol/l and 98-2,863 pmol/l.
“Metabolitelevels of progesterone, as measured in the skin secretions of two females for six consecutive months, were 104-2,662 pmol/l and 98-2,863 pmol/l (Dathe and Kuckelkorn 1989).”
Quote in the paperSubstantial claim0 of 1 sources checked- cites Dathe and Kuckelkorn 1989
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Metabolite concentrations of estrogen in fecal samples indicated a mean (± SD) estrous cycle of 31.8 ± 7.4 days.
“Metabolite concentrations of estrogen in fecal samples similarly indicated a mean (± SD) estrous cycle of 31.8 ± 7.4 days (Flacke et al. 2017a).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Flacke et al. 2017a
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Direct and indirect observations repeatedly provide evidence that C. liberiensis is predominantly solitary except during the breeding season, or as a mother and calf pair.
“Direct and indirect (camera trap photos, footprint tracking) observations repeatedly provide evidence that C. liberiensis is predominantly solitary except during the breeding season, or as a mother and calf pair (Schomburgk 1922;Robinson 1970;Bülow 1987;Hentschel 1990;Collen et al. 2011;Eshuis 2011;van Heukelum 2011;Conway 2013).”
Quote in the paperNot a substantial claim0 of 3 sources checked- cites Schomburgk 1922
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Robinson 1970
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Bülow 1987
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
At the end of 2016, the captive population of C. liberiensis was comprised of 153 males, 229 females, and nine animals of undetermined sex living in 144 zoological institutions and private collections worldwide.
“At the end of 2016, the captive population of C. liberiensis was comprised of 153 males, 229 females, and nine animals of undetermined sex living in 144 zoological institutions and private collections worldwide (Steck 2017).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Steck 2017
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
The captive population is female-biased: 40.1% males and 59.9% females for animals of known sex.
“The captive population is female-biased: 40.1% males and 59.9% females for animals of known sex (Steck 2017).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Steck 2017
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisecitation
Abundance of C. liberiensis has an inverse relationship to human population density and intensity of land use.
“Abundance of C. liberiensis has an inverse relationship to human population density and intensity of land use (Robinson 1970).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Robinson 1970
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Premisenone given
This captive population is derived from 162 founders from these countries: Liberia-89; Côte d'Ivoire-23; Sierra Leone-8; Unknown-42.
“This population is derived from 162 founders from these countries: Liberia-89; Côte d'Ivoire-23; Sierra Leone-8; Unknown-42.”
Quote in the paperNot a substantial claim - Premisecitation
Field survey data and community questionnaires conducted in the Gola transboundary region showed that C. liberiensis is frequently using areas of unprotected farm bush.
“Field survey data (transect walks and camera traps) and community questionnaires conducted in the Gola transboundary region between Sierra Leone and Liberia showed that C. liberiensis is frequently using areas of unprotected farm bush; thus, a mixed conservation approach of expanding both protected area networks together with community-based conservation activities is necessary to ensure long-term”
Quote in the paperSubstantial claim0 of 1 sources checked- cites Hillers et al. 2016
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- Classified as probably not a substantial claim. 42
- May combine two claims. 92
- The quote only partly states this claim. 1
Probably not a substantial claim: may not pass S3 (load-bearing).
PremisecitationRarely there are three total mandibular incisors instead of the single pair in C. liberiensis.
“Rarely there are three total mandibular incisors instead of the single pair (Johnston 1906).”
Quote in the paperNot a substantial claim1 of 1 sources checked- cites Johnston 1906 source
The available copy’s passages on Liberian fauna, including the discussion near p. 89, do not address the number of mandibular incisors in C. liberiensis. Searches for the species, pygmy hippopotamus, and incisors found no relevant passage; the copy is truncated, so the decisive description may be missing. (LLM, conf=0.73)
Experimental — accuracy not yet measured
Probably not a substantial claim: may not pass S3 (load-bearing).
PremisecitationRecent molecular phylogenetic studies and comparative morphological analyses support Hippopotamidae and Cetacea as forming a monophyletic group.
“Recent molecular phylogenetic studies and comparative morphological analyses have supported Hippopotamidae and Cetacea as forming a monophyletic group (Shimamura et al. 1997;Nomura et al. 1998;Ursing and Arnason 1998;Nomura and Yasue 1999;Geisler and Uhen 2003;Boisserie 2005;Boisserie et al. 2005;Fisher et al. 2007);”
Quote in the paperNot a substantial claim2 of 3 sources checked- cites Shimamura et al. 1997 source
- cites Boisserie 2005 source
- cites Fisher et al. 2007 source
The Abstract says molecular studies indicate that hippos and cetaceans are sister taxa. In “Functional and Phylogenetic Considerations,” the authors say their forelimb comparisons support the molecular data, but they call for further study to determine whether hippos were closely allied to Cetacea. The paper therefore supports the molecular part of the claim, but does not establish that its comparative morphological analysis supports the hippo–cetacean clade. (LLM, conf=0.87)
Experimental — accuracy not yet measured
May combine two claims.
PremisecitationProportionally, the brain case is larger and the muzzle is shorter in C. liberiensis than in the common hippopotamus.
“Proportionally, the brain case is larger and the muzzle is shorter in C. liberiensis than in the common hippopotamus (Fig. 2; Johnston 1906).”
Quote in the paperNot a substantial claim1 of 1 sources checked- cites Johnston 1906 source
The passage near p. 89 discusses Liberian animals but does not compare the skulls of C. liberiensis and the common hippopotamus. Searches for the species name, brain case, muzzle, and skull comparison found no relevant passage in our copy, whose end may be missing. (LLM, conf=0.74)
Experimental — accuracy not yet measured
Classified as probably not a substantial claim. May combine two claims.
PremisecitationBoisserie (2005) concluded that the mix of primitive and derived features of the extant pygmy hippo gives it a distinct lineage, validating the genus Choeropsis for the pygmy hippo and restricting the genus Hexaprotodon to the fossil lineage found mostly in Asia.
“Most recently, Boisserie (2005) concluded that the mix of primitive and derived features of the extant pygmy hippo gives it a distinct lineage, validating the genus Choeropsis for the pygmy hippo and restricting the genus Hexaprotodon to the fossil lineage found mostly in Asia (Falconer and Cautley 1836).”
Quote in the paperSubstantial claim1 of 2 sources checked- cites Boisserie 2005 source
- cites Falconer and Cautley 1836
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
InterpretationcitationOverall, the myology of C. liberiensis resembles ruminants more closely than suids, and both extant hippo species are unique in retaining several primitive features in the forelimb, consistent with the hypothesis that hippopotamids diverged from other Artiodactyla early in the history of this group.
“Overall, the myology of the C. liberiensis resembles ruminants more closely than suids, and both extant hippo species are unique in retaining several primitive features in the forelimb, consistent with the hypothesis that hippopotamids diverged from other Artiodactyla early in the history of this group (Fisher et al. 2007).”
Quote in the paperSubstantial claim1 of 1 sources checked- cites Fisher et al. 2007 source
May combine two claims.
PremisecitationOther recent confirmed records of C. liberiensis are from Gola National Forest in the west, from the Wonegizi National Forest in the north, on the border with Guinea, and in Sapo National Park in the southeast.
“Other recent confirmed records are from Gola National Forest in the west, from the Wonegizi National Forest in the north, on the border with Guinea, and in Sapo National Park in the southeast (Collen et al. 2011).”
Quote in the paperNot a substantial claim1 of 1 sources checked- cites Collen et al. 2011 source
The Abstract reports seven camera-trap photographs confirming C. liberiensis in Sapo National Park, Liberia. Our copy does not report confirmed records from Gola National Forest or Wonegizi National Forest, so the citation supports only the Sapo portion of the sentence. (LLM, conf=0.88)
Experimental — accuracy not yet measured
May combine two claims.
PremisecitationThe insertion site of the brachialis muscle can be on the radius only or on both the radius and ulna in C. liberiensis.
“The insertion site of the brachialis muscle can be on the radius only (Macalister 1873;Fisher et al. 2007) or on both the radius and ulna (Campbell 1936); the radius and ulna are functionally or actually fused in artiodactyls.”
Quote in the paperNot a substantial claim1 of 3 sources checkedProbably not a substantial claim: may not pass S3 (load-bearing).
PremisecitationHippopotamids are believed to have originated in Africa sometime in the mid-Miocene period, approximately 11-16 million years ago, in the areas of modern-day Kenya and Tunisia.
“Hippopotamids are believed to have originated in Africa sometime in the mid-Miocene period, approximately 11-16 million years ago, in the areas of modern-day Kenya and Tunisia (Pickford 1983;Boisserie 2005).”
Quote in the paperNot a substantial claim1 of 2 sources checked- cites Pickford 1983 source
- cites Boisserie 2005 source
The Introduction says Hippopotamidae most probably had their roots in Africa during the middle Miocene. It mentions a Miocene hippopotamid from Kenya, but does not identify Kenya or Tunisia as places of origin or give the specific 11–16-million-year range. (LLM, conf=0.87)
Experimental — accuracy not yet measured
May combine two claims.
PremisecitationC. liberiensis has retained several features more adapted to a terrestrial lifestyle compared to the common hippopotamus, including proportionally longer limbs and only moderately webbed toes that spread more widely under its own weight.
“Choeropsis liberiensis has retained several features more adapted to a terrestrial lifestyle compared to the common hippopotamus, including proportionally longer limbs and only moderately webbed toes that spread more widely under its own weight (Johnston 1906 ;Pocock 1923;Fisher et al. 2007).”
Quote in the paperNot a substantial claim2 of 3 sources checked- cites Johnston 1906 source
The available copy’s passages on Liberian fauna (Chapter II) do not describe Choeropsis liberiensis or compare its limbs and toes with those of the common hippopotamus. Searches for hippopotamus, webbed toes, and the claimed limb proportions found no relevant passage; the copy is truncated, so the relevant discussion may be missing. (LLM, conf=0.72)
Experimental — accuracy not yet measured - cites Pocock 1923 source
- cites Fisher et al. 2007 source
The Functional and Phylogenetic Considerations section says hippos retain terrestrial locomotor patterns and have tendons that control toe splay on soft ground. It does not establish that pygmy hippos have proportionally longer limbs, moderately webbed toes, or greater terrestrial adaptation than common hippos; the Abstract says common hippos share the described toe-control features. (LLM, conf=0.85)
Experimental — accuracy not yet measured
May combine two claims.
PremisecitationThe trapezius muscle can variably originate from the last two cervical vertebrae or the occipital bone in C. liberiensis, whereas the common hippopotamus lacks an occipital origin.
“Similarly, the trapezius muscle can variably originate from the last two cervical vertebrae (Campbell 1936) or the occipital bone (Macalister 1873;Fisher et al. 2007) in C. liberiensis, whereas the common hippopotamus lacks an occipital origin.”
Quote in the paperNot a substantial claim1 of 3 sources checkedProbably not a substantial claim: may not pass S3 (load-bearing). May combine two claims.
PremisecitationThe Cetacea and the Hippopotamidae are believed to share a common ancestry, with initial divergence estimated around 54 million years ago.
“The Cetacea and the Hippopotamidae are believed to share a common ancestry, with initial divergence estimated around 54 million years ago (Ursing and Arnason 1998).”
Quote in the paperNot a substantial claim1 of 1 sources checked- cites Ursing and Arnason 1998 source
May combine two claims.
PremisecitationThe current estimate for Sierra Leone is roughly 150 C. liberiensis individuals countrywide, primarily within the Gola Forest complex.
“Data from recent field surveys were not sufficient to determine actual abundance (Conway 2013); the current estimate for Sierra Leone is roughly 150 individuals countrywide, primarily within the Gola Forest complex, including the Moa River islands (Lindsell et al. 2011;Mallon et al. 2011).”
Quote in the paperNot a substantial claim1 of 3 sources checked- cites Conway 2013
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Lindsell et al. 2011 source
The Results (page not provided) document pygmy hippopotamuses in the Gola area and on Tiwai Island in the Moa River. The Discussion (page not provided) suggests several tens of animals in and around the reserves, but does not give a countrywide estimate of roughly 150 or establish that most of Sierra Leone’s population is there. (LLM, conf=0.88)
Experimental — accuracy not yet measured - cites Mallon et al. 2011
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationPopulation surveys conducted in Liberia have indicated a minimum of 1,000 C. liberiensis and perhaps several thousand.
“Population surveys conducted in Liberia have indicated a minimum of 1,000 (Verschuren 1982) and perhaps several thousand (Eltringham 1993).”
Quote in the paperNot a substantial claim1 of 2 sources checked- cites Verschuren 1982 source
- cites Eltringham 1993
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationFurther threats to C. liberiensis include lack of adequate financial and legal protection for the few remaining intact areas of habitat, poaching for bushmeat, and increasing human settlement adjacent to protected areas.
“Further threats include lack of adequate financial and legal protection for the few remaining intact areas of habitat, poaching for bushmeat, and increasing human settlement adjacent to protected areas (Anstey 1991b;Critical Ecosystem Partnership Fund 2015;Lindsell et al. 2011;Ransom et al. 2015).”
Quote in the paperNot a substantial claim1 of 3 sources checked- cites Anstey 1991b
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Critical Ecosystem Partnership Fund 2015
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Lindsell et al. 2011 source
The Discussion (page unavailable) says that 63% of riverine forest in the Gola area lies outside the reserves, so pygmy hippopotamuses may not be well protected. It also identifies an increasing human population and demand for bushmeat as threats to the forest. It does not establish inadequate financial or legal protection, or identify poaching and settlement adjacent to protected areas as specific threats to C. liberiensis. (LLM, conf=0.84)
Experimental — accuracy not yet measured
Probably not a substantial claim: may not pass S3 (load-bearing).
Findingown dataThe greatest length of the skull of an adult female Choeropsis liberiensis specimen (ZFMK MAM 1965.0570) from Côte d'Ivoire is 339 mm.
“Greatest length of skull is 339 mm.”
Quote in the paperNot a substantial claimProbably not a substantial claim: may not pass S3 (load-bearing).
InterpretationcitationThe animals at the higher end of the weight range are undoubtedly overweight.
“The animals at the higher end of the weight range are undoubtedly overweight (Flacke et al. 2015).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Flacke et al. 2015 source
Probably not a substantial claim: may not pass S3 (load-bearing).
PremisecitationThe view validating Choeropsis was also recently embraced by Groves and Grubb (2011).
“This view was also recently embraced by Groves and Grubb (2011).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Groves and Grubb 2011
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationAverage weight in captivity for C. liberiensis ranges from 160 to 275 kg with a shoulder height of 70-80 cm.
“Average weight in captivity ranges from 160 to 275 kg with a shoulder height of 70-80 cm (Lang 1975);”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Lang 1975
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
Probably not a substantial claim: may not pass S3 (load-bearing).
Premisenone givenThe common hippopotamus frequents riverine savannahs.
“C. liberiensis inhabits primary and secondary rainforest and swamps (Bülow 1987;Hentschel 1990), whereas the common hippopotamus frequents riverine savannahs.”
Quote in the paperNot a substantial claimMay combine two claims.
PremisereasoningC. liberiensis is distinguished from the common hippopotamus Hippopotamus amphibius by its smaller size, dental formula, distribution, and habitat.
“Choeropsis liberiensis (Fig. 1) is distinguished from the common hippopotamus Hippopotamus amphibius by its smaller size, dental formula, distribution, and habitat.”
Quote in the paperNot a substantial claimProbably not a substantial claim: may not pass S3 (load-bearing).
Premisenone givenC. liberiensis is the smaller of the two extant hippopotamids.
“The smaller of the two extant hippopotamids, it is relatively uncommon in zoological facilities and private collections worldwide.”
Quote in the paperNot a substantial claimProbably not a substantial claim: may not pass S3 (load-bearing).
PremisecitationThe common hippopotamus is found throughout sub-Saharan Africa.
“the common hippopotamus is found throughout sub-Saharan Africa (Boisserie et al. 2005).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Boisserie et al. 2005 source
Probably not a substantial claim: may not pass S3 (load-bearing).
Premisenone givenC. liberiensis is gray to black in color with occasional depigmented pink patches on the body and limbs.
“Choeropsis liberiensis is gray to black in color with occasional depigmented pink patches on the body and limbs (Fig. 1).”
Quote in the paperNot a substantial claimProbably not a substantial claim: may not pass S3 (load-bearing).
PremisecitationCoarse, bristle-like hairs are found only at the ear margins, on the lips, and at the end of the tail in C. liberiensis.
“Coarse, bristle-like hairs are found only at the ear margins, on the lips, and at the end of the tail (Lochte 1951).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Lochte 1951
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
Probably not a substantial claim: may not pass S3 (load-bearing). May combine two claims.
PremisecitationA wild common hippopotamus can weigh upwards of 1,500 kg with a shoulder height of 130-140 cm.
“by comparison, a wild common hippopotamus can weigh upwards of 1,500 kg with a shoulder height of 130-140 cm (Marshall and Sayer 1976).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Marshall and Sayer 1976 source
Probably not a substantial claim: may not pass S3 (load-bearing).
Premisenone givenC. liberiensis is a CITES Appendix II species.
“A solitary, forest-dwelling ungulate endemic to the Upper Guinean forest ecosystem in West Africa, it is classified as “Endangered” by the International Union for Conservation of Nature and Natural Resources and is a Convention on International Trade in Endangered Species of Wild Flora and Fauna (CITES) Appendix II species.”
Quote in the paperNot a substantial claimMay combine two claims.
PremisecitationC. liberiensis is limited in geographic distribution to the Upper Guinean forest ecosystem of West Africa and is now found only in Côte d'Ivoire, Guinea, Liberia, and Sierra Leone.
“C. liberiensis is limited in geographic distribution to the Upper Guinean forest ecosystem of West Africa and is now found only in Côte d'Ivoire, Guinea, Liberia, and Sierra Leone (Ransom et al. 2015);”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Ransom et al. 2015
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
Probably not a substantial claim: may not pass S3 (load-bearing), S4 (based).
Premisenone givenC. liberiensis is relatively uncommon in zoological facilities and private collections worldwide.
“The smaller of the two extant hippopotamids, it is relatively uncommon in zoological facilities and private collections worldwide.”
Quote in the paperNot a substantial claimProbably not a substantial claim: may not pass S3 (load-bearing).
Premisenone givenC. liberiensis is classified as Endangered by the International Union for Conservation of Nature and Natural Resources.
“A solitary, forest-dwelling ungulate endemic to the Upper Guinean forest ecosystem in West Africa, it is classified as “Endangered” by the International Union for Conservation of Nature and Natural Resources and is a Convention on International Trade in Endangered Species of Wild Flora and Fauna (CITES) Appendix II species.”
Quote in the paperNot a substantial claimMay combine two claims.
PremisecitationThe International Studbook reports a body mass range of 180-260 kg and a shoulder height of 80 cm for C. liberiensis.
“The International Studbook reports a body mass range of 180-260 kg and a shoulder height of 80 cm (Steck 2017);”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Steck 2017
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
Probably not a substantial claim: may not pass S3 (load-bearing). May combine two claims.
Premisenone givenC. liberiensis is a solitary, forest-dwelling ungulate endemic to the Upper Guinean forest ecosystem in West Africa.
“A solitary, forest-dwelling ungulate endemic to the Upper Guinean forest ecosystem in West Africa, it is classified as “Endangered” by the International Union for Conservation of Nature and Natural Resources and is a Convention on International Trade in Endangered Species of Wild Flora and Fauna (CITES) Appendix II species.”
Quote in the paperNot a substantial claimProbably not a substantial claim: may not pass S3 (load-bearing).
PremisecitationSubdermal glands of C. liberiensis secrete an oily clear-to-white, foamy fluid that protects the skin against sunburn, dehydration, and bacterial infection.
“Subdermal glands secrete an oily clear-to-white, foamy fluid that protects the skin against sunburn, dehydration, and bacterial infection (Eulenberger 1995;Hashimoto et al. 2007).”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Eulenberger 1995
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Hashimoto et al. 2007 source
Probably not a substantial claim: may not pass S3 (load-bearing). May combine two claims.
PremisecitationThe skin of C. liberiensis is thick with a rubbery texture.
“The skin is thick with a rubbery texture (Flach et al. 1998).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Flach et al. 1998 source
May combine two claims.
PremisecitationBody mass for adult female C. liberiensis (n = 5) ranged from 179 to 267 kg; for adult males (n = 2) 208 kg and 273 kg.
“Body mass for adult females (n = 5) ranged from 179 to 267 kg; for adult males (n = 2) 208 kg and 273 kg (Lang 1975).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Lang 1975
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
Premisenone givenThere is a dearth of detailed information concerning Choeropsis in the wild due to the limitations for in situ research; the majority of data is derived from zoo specimens, all of which are of the nominate species C. liberiensis.
“There is a dearth of detailed information concerning Choeropsis in the wild due to the limitations for in situ research; the majority of data is derived from zoo specimens, all of which are of the nominate species C. liberiensis.”
Quote in the paperNot a substantial claimMay combine two claims.
PremisecitationThe extant Choeropsis liberiensis is endemic only to the Upper Guinean forest ecosystem in West Africa, and its distribution is limited to fragmented populations in Côte d'Ivoire, Guinea, Liberia, and Sierra Leone.
“The extant Choeropsis liberiensis is endemic only to the Upper Guinean forest ecosystem in West Africa (Fig. 3), and its distribution is limited to fragmented populations in Côte d'Ivoire, Guinea, Liberia, and Sierra Leone (Ransom et al. 2015).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Ransom et al. 2015
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
Probably not a substantial claim: may not pass S3 (load-bearing).
PremisecitationCetartiodactyla was proposed as the taxon name for the group formed by Hippopotamidae and Cetacea.
“Cetartiodactyla was proposed as the taxon name for the group (Montgelard et al. 1997) and subsequently more clearly defined (O' Leary and Gatesy 2008).”
Quote in the paperNot a substantial claim0 of 2 sources checkedProbably not a substantial claim: may not pass S3 (load-bearing).
PremisecitationThe ears and nostrils of C. liberiensis function as valves that close tightly when the animal is under water.
“The ears and nostrils function as valves that close tightly when the animal is under water (Pocock 1923).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Pocock 1923 source
May combine two claims.
PremisecitationThe four stomach compartments of C. liberiensis are similar to those of a ruminant and include the nonglandular visceral and parietal blind sacs (also termed left and right diverticula), the vestibulum, the connecting chamber, and the customary glandular stomach.
“The four stomach compartments are similar to those of a ruminant and include the nonglandular visceral and parietal blind sacs (also termed left and right diverticula), the vestibulum, the connecting chamber, and the customary glandular stomach (Langer 1975;Macdonald and Hartman 1983).”
Quote in the paperNot a substantial claim0 of 2 sources checkedMay combine two claims.
PremisecitationThe highly vascularized and inelastic skin of C. liberiensis is about 1 cm thick with extensive subcutaneous fat and a rubbery texture.
“The highly vascularized and inelastic skin of C. liberiensis is about 1 cm thick with extensive subcutaneous fat and a rubbery texture (Flach et al. 1998).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Flach et al. 1998 source
May combine two claims.
PremisecitationThe volume of the combined fore stomachs in the newborn C. liberiensis is about 68% of the total gastric volume; in adults this proportion increases to 94%.
“The volume of the combined fore stomachs in the newborn C. liberiensis is about 68% of the total gastric volume; in adults this proportion increases to 94% (Langer 1975).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Langer 1975 source
Probably not a substantial claim: may not pass S3 (load-bearing). May combine two claims.
PremisecitationThe vertebral formula of C. liberiensis is 7C, 14T, 5L, 4S, and 10 or more CA, total 40 or more.
“The vertebral formula of C. liberiensis is 7C, 14T, 5L, 4S, and 10 or more CA, total 40 or more (Macalister 1873).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Macalister 1873
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationSkin secretions from a female C. liberiensis had a pH of 9.5 and lower mean sodium (Na+) and potassium (K+) concentrations than secretions from the common hippopotamus, possibly reflecting dietary or metabolic differences.
“Skin secretions from a female C. liberiensis had a pH of 9.5 and lower mean sodium (Na + ) and potassium (K + ) concentrations than secretions from the common hippopotamus, possibly reflecting dietary or metabolic differences (Olivier 1975).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Olivier 1975 source
May combine two claims.
PremisecitationConfirmed reports of C. liberiensis presence are from both Tiwai Island and other small islands along the Moa River, and from the Gola Forest Reserve Complex along the Mano River on the border with Liberia.
“Confirmed reports of its presence are from both Tiwai Island and other small islands along the Moa River (White et al. 1986;Conway 2013), and from the Gola Forest Reserve Complex along the Mano River on the border with Liberia (Hillers and Muana 2010;Lindsell et al. 2011;Mallon et al. 2011).”
Quote in the paperNot a substantial claim0 of 3 sources checked- cites White et al. 1986
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Conway 2013
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Mallon et al. 2011
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationThe connecting chamber is oriented in a dorsal-to-ventral direction in the newborn to assist with passage of milk into the glandular stomach; the orientation changes in the adult to promote retention of ingesta and its movement between compartments of the fore stomach.
“The connecting chamber is oriented in a dorsal-toventral direction in the newborn to assist with passage of milk into the glandular stomach; the orientation changes in the adult to promote retention of ingesta and its movement between compartments of the fore stomach (Macdonald and Hartman 1983).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Macdonald and Hartman 1983 source
May combine two claims.
PremisecitationInstead of infundibula and renculi, the lobes of C. liberiensis kidneys are projections like the fingers of a hand; each kidney has 17-25 lobes.
“Instead, the lobes are projections like the fingers of a hand; each kidney has 17-25 lobes (Maluf 1978(Maluf , 1994)).”
Quote in the paperNot a substantial claim0 of 2 sources checkedMay combine two claims.
PremisecitationThe globe (eyeball) of C. liberiensis is rounded and slightly taller than it is wide; measurements from a subadult female (117 kg) were 24.3 mm horizontal diameter, 18.6 mm anterior to posterior diameter, and 22.5 mm vertical diameter.
“The globe (eyeball) itself is rounded and slightly taller than it is wide; measurements from a subadult female (117 kg) were 24.3 mm horizontal diameter, 18.6 mm anterior to posterior diameter, and 22.5 mm vertical diameter (Hegner 1967).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Hegner 1967 source
May combine two claims.
PremisecitationThe species has also been confirmed in the southeast in Maryland and River Gee counties and within the proposed Grand Kru-River Gee Protected Area.
“The species has also been confirmed in the southeast in Maryland and River Gee counties and within the proposed Grand Kru-River Gee Protected Area (Mallon et al. 2011).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Mallon et al. 2011
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationThe tapetum lucidum of C. liberiensis is light blue, reflective, and extends across the majority of the fundus; the ventral aspect is grossly obscured by the pigmented epithelial cell layer of the retina and thus the functional tapetum primarily occupies the dorsal aspect of the fundus.
“The tapetum lucidum is light blue, reflective, and extends across the majority of the fundus; the ventral aspect is grossly obscured by the pigmented epithelial cell layer of the retina and thus the functional tapetum primarily occupies the dorsal aspect of the fundus (Hegner 1967).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Hegner 1967 source
May combine two claims.
InterpretationcitationC. liberiensis has low-crowned molars, further supporting its status as a forest-dwelling consumer of coarse vegetation.
“C. liberiensis has low-crowned molars, further supporting its status as a forest-dwelling consumer of coarse vegetation (Weston 2000).”
Quote in the paperSubstantial claim0 of 1 sources checked- cites Weston 2000 source
May combine two claims.
PremisecitationThe entire small intestine (duodenum, jejunum, ileum) of an adult female C. liberiensis is about 16 m long with an average diameter of 2.5 cm; the colon is about 2.5 m long.
“The entire small intestine (duodenum, jejunum, ileum) of an adult female is about 16 m long with an average diameter of 2.5 cm; the colon is about 2.5 m long (Macdonald and Hartman 1983).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Macdonald and Hartman 1983 source
May combine two claims.
Premisenone givenThe eyes of C. liberiensis are less prominent and have a more lateral placement with a wider field of vision than with the dorsal eye placement of the amphibious common hippopotamus.
“The eyes of C. liberiensis are less prominent and have a more lateral placement with a wider field of vision than with the dorsal eye placement of the amphibious common hippopotamus.”
Quote in the paperNot a substantial claimProbably not a substantial claim: may not pass S3 (load-bearing), S4 (based).
Premisenone givenSimilar conglomerate, bristle-like hairs are found only in seals and sea lions (Carnivora; Otariidae).
“Similar conglomerate, bristle-like hairs are found only in seals and sea lions (Carnivora; Otariidae).”
Quote in the paperNot a substantial claimProbably not a substantial claim: may not pass S3 (load-bearing). May combine two claims.
PremisecitationHair measurements (mm) for C. liberiensis were: up to 35 for lip hairs, 6-9 for ear hairs, and 10-18 for tail hairs.
“Hair measurements (mm) for C. liberiensis were: up to 35 for lip hairs, 6-9 for ear hairs, and 10-18 for tail hairs (Lochte 1951).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Lochte 1951
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
Probably not a substantial claim: may not pass S3 (load-bearing).
Premisenone givenFossil records support the much more recent presence of the now-extinct dwarf hippo species on the islands of Crete (Hippopotamus creutzburgi), Cyprus (Phanourios minor), and Madagascar (Hippopotamus madagascariensis).
“Fossil records support the much more recent presence of the now-extinct dwarf hippo species on the islands of Crete (Hippopotamus creutzburgi), Cyprus (Phanourios minor), and Madagascar (Hippopotamus madagascariensis).”
Quote in the paperNot a substantial claimMay combine two claims.
PremisecitationThe retina of C. liberiensis has two histologically distinct types of pigmented epithelial cells and three types of photoreceptor cells.
“The retina itself has two histologically distinct types of pigmented epithelial cells and three types of photoreceptor cells (Hegner 1967).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Hegner 1967 source
May combine two claims.
PremisecitationEach kidney of C. liberiensis has about three times 10^6 glomeruli; however, the total length of the tubule, including the Loop of Henle, is only 6.8 mm.
“Each kidney has about three times 10 6 glomeruli (Maluf 1994); however, the total length of the tubule, including the Loop of Henle, is only 6.8 mm (Maluf 1978).”
Quote in the paperSubstantial claim0 of 2 sources checkedProbably not a substantial claim: may not pass S3 (load-bearing).
PremisecitationThe earliest Hexaprotodon fossils, dating from the Mio-Pliocene period about 5 million years ago, were recovered from the Siwalik Hills on the India-Pakistan border.
“The earliest Hexaprotodon fossils, dating from the Mio-Pliocene period about 5 million years ago, were recovered from the Siwalik Hills on the India-Pakistan border (Falconer and Cautley 1836).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Falconer and Cautley 1836
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationMultiple anecdotal reports describe white foamy skin secretions produced by the subdermal glands in wild C. liberiensis, and captive animals often exhibit the same secretions during periods of physical exertion, including during mating.
“Multiple anecdotal reports describe white foamy skin secretions produced by the subdermal glands in wild C. liberiensis, and captive animals often exhibit the same secretions during periods of physical exertion, including during mating (Steinmetz 1937;Lang 1975;Schubert 2004;von Houwald et al. 2007).”
Quote in the paperNot a substantial claim0 of 3 sources checked- cites Steinmetz 1937
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Lang 1975
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites von Houwald et al. 2007
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationAn elongated triangular gall bladder is present in C. liberiensis, and the bile duct empties into the duodenum 15 cm distal to the pylorus.
“An elongated triangular gall bladder is present, and the bile duct empties into the duodenum 15 cm distal to the pylorus (Macalister 1873;Langer 1975).”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Macalister 1873
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Langer 1975 source
Probably not a substantial claim: may not pass S3 (load-bearing).
PremisecitationOne animal was caught in a pit-trap near Tiwai Island in 2010.
“One animal was also caught in a pit-trap near Tiwai Island in 2010 (Conway 2013).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Conway 2013
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
InterpretationcitationThis range is comparable to previously published rates for the common hippopotamus at the same temperatures, indicative of similar dermal physiology for the two species.
“This range is comparable to previously published rates for the common hippopotamus at the same temperatures, indicative of similar dermal physiology for the two species (Olivier 1975).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Olivier 1975 source
May combine two claims.
Premisenone givenField surveys conducted between 1982 and 1986 found signs of C. liberiensis within 21 protected areas in southwestern Côte d'Ivoire, with the highest abundances in Taï National Park and the surrounding N'Zo Faunal Reserve.
“Field surveys conducted between 1982 and 1986 found signs of C. liberiensis within 21 protected areas in southwestern Côte d'Ivoire, with the highest abundances in Taï National Park and the surrounding N'Zo Faunal Reserve.”
Quote in the paperNot a substantial claimMay combine two claims.
PremisecitationBy the late 20th century, the distribution of C. liberiensis was limited to Sapo National Park and Grebo National Forest in the southeast, and North Lorma National Forest in the far northwest; it was deemed unlikely to persist in the central regions of Liberia.
“However, by the late 20th century its distribution was limited to Sapo National Park and Grebo National Forest in the southeast, and North Lorma National Forest in the far northwest; it was deemed unlikely to persist in the central regions of Liberia (Anstey 1991a).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Anstey 1991a
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationThe two kidneys equal 0.22% of body mass in the adult and 0.51-0.63% of body mass in the newborn C. liberiensis.
“The two kidneys equal 0.22% of body mass in the adult and 0.51-0.63% of body mass in the newborn (Maluf 1994).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Maluf 1994 source
May combine two claims.
PremisecitationA 2005 survey conducted in Gola, North Lorma, and Grebo National Forests in Liberia confirmed signs of C. liberiensis only in Grebo National Forest, in the far southeast near the border with Côte d'Ivoire; the other areas lacked evidence of its presence.
“A 2005 survey conducted in Gola, North Lorma, and Grebo National Forests in Liberia confirmed signs of C. liberiensis only in Grebo National Forest, in the far southeast near the border with Côte d'Ivoire; the other areas lacked evidence of its presence (Barrie et al. 2007).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Barrie et al. 2007 source
May combine two claims.
PremisecitationThe mucosal epithelium of the fore stomachs is lined with finger-like papillae, each varying in length, thickness, and shape; these papillae are less prominent in the newborn.
“The mucosal epithelium of the fore stomachs is lined with finger-like papillae, each varying in length, thickness, and shape; these papillae are less prominent in the newborn (Macdonald and Hartman 1983;Endo et al. 2001).”
Quote in the paperNot a substantial claim0 of 2 sources checkedMay combine two claims.
InterpretationcitationThe most westward extent of the range of C. liberiensis is likely the Ouatamba-Kilimi National Park in northwestern Sierra Leone, also the northernmost location where the species has been recorded, although there are no recent reports to substantiate its continued presence in this area.
“The most westward extent of its range is likely the Ouatamba-Kilimi National Park in northwestern Sierra Leone (White 1986), also the northernmost location where the species has been recorded, although there are no recent reports to substantiate its continued presence in this area.”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites White 1986
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationIn Ziama, evidence of C. liberiensis (tracks and dung) was found along the Njéré, Gniogné, and upper Ouin watercourses, and in Diécké along the Nié, Grand Gbin, Gbin-bé, and Lih streams.
“In Ziama, evidence of C. liberiensis (tracks and dung) was found along the Njéré, Gniogné, and upper Ouin watercourses, and in Diécké along the Nié, Grand Gbin, Gbin-bé, and Lih streams (Bützler 1994).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Bützler 1994
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationThere is no cecum in C. liberiensis, but the junction between the large and small intestine is characterized by an increase in diameter and a change in the lining of the mucosal epithelium.
“There is no cecum (Macalister 1873), but the junction between the large and small intestine is characterized by an increase in diameter and a change in the lining of the mucosal epithelium (Macdonald and Hartman 1983).”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Macalister 1873
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Macdonald and Hartman 1983 source
May combine two claims.
PremisecitationLobulated kidneys of C. liberiensis are similar in architecture to those of the Bovidae but lack infundibula and renculi.
“Lobulated kidneys are similar in architecture to those of the Bovidae but lack infundibula and renculi (Macalister 1873).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Macalister 1873
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
Probably not a substantial claim: may not pass S3 (load-bearing), S4 (based). May combine two claims.
Premisenone givenThe hippopotamids are unique among the Artiodactyla in that they have retained a semiaquatic lifestyle and all four toes are weight-bearing.
“The hippopotamids are unique among the Artiodactyla in that they have retained a semiaquatic lifestyle and all four toes are weight-bearing.”
Quote in the paperNot a substantial claimMay combine two claims.
PremisecitationThe pupil of C. liberiensis is oval, 7 mm wide and 5 mm high, with a symmetrical iris width of 2 mm; the lens is 10 mm in diameter and 7 mm thick; scleral thickness varies between 1.2 and 1.5 mm.
“The pupil is oval, 7 mm wide and 5 mm high, with a symmetrical iris width of 2 mm; the lens is 10 mm in diameter and 7 mm thick; scleral thickness varies between 1.2 and 1.5 mm (Hegner 1967).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Hegner 1967 source
May combine two claims.
PremisecitationPolycystic kidney disease, first noted as an incidental finding at necropsy, is now recognized as a possible X-linked dominant inheritance trait, with prevalence rates greater than 37% in captive animals.
“Polycystic kidney disease, first noted as an incidental finding at necropsy (Jarofke and Klös 1982), is now recognized as a possible X-linked dominant inheritance trait, with prevalence rates greater than 37% in captive animals (Raymond et al. 2000;Nees et al. 2009;Flacke et al. 2017b).”
Quote in the paperNot a substantial claim0 of 3 sources checked- cites Jarofke and Klös 1982
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Raymond et al. 2000
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Nees et al. 2009 source
May combine two claims.
PremisecitationCommunal housing of C. liberiensis can lead to conspecific aggression resulting in injury and death, may contribute to abnormal behaviors including stereotypies, and is also believed to compromise normal maternal behavior in some circumstances.
“Communal housing can lead to conspecific aggression resulting in injury and death, may contribute to abnormal behaviors including stereotypies, and is also believed to compromise normal maternal behavior in some circumstances (Flacke et al. 2015).”
Quote in the paperSubstantial claim0 of 1 sources checked- cites Flacke et al. 2015 source
May combine two claims.
PremisecitationC. liberiensis is found in both primary and secondary rainforests; low-lying Raphia palm swamps are its preferred habitat type because these offer both feeding and resting areas.
“C. liberiensis is found in both primary and secondary rainforests; low-lying Raphia palm swamps are its preferred habitat type because these offer both feeding and resting areas (Robinson 1970;Bülow 1987).”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Robinson 1970
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Bülow 1987
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisereasoningThese measures of genetic diversity are calculated via analysis of studbook data, data that inaccurately assume equal relatedness among founders from wild populations, may contain errors in recorded pedigrees, and cannot clearly identify genetically underrepresented animals.
“However, these measures of genetic diversity are calculated via analysis of studbook data, data that inaccurately assume equal relatedness among founders from wild populations, may contain errors in recorded pedigrees, and cannot clearly identify genetically underrepresented animals.”
Quote in the paperSubstantial claimMay combine two claims.
PremisecitationRecently, 1.2 mg/kg ketamine and 0.08 mg/kg medetomidine followed by endotracheal intubation and maintenance on oxygen and isoflurane gas has proven very safe and effective for anesthesia of C. liberiensis, with no reported mortality.
“Recently, 1.2 mg/kg ketamine and 0.08 mg/kg medetomidine followed by endotracheal intubation and maintenance on oxygen and isoflurane gas has proven very safe and effective for anesthesia, with no reported mortality (Bouts et al. 2012).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Bouts et al. 2012 source
May combine two claims.
PremisecitationVery young C. liberiensis calves are hidden in a shallow pool or among the vegetation while the mother forages; she will visit the calf several times per day to allow it to nurse.
“Very young calves are hidden in a shallow pool or among the vegetation while the mother forages; she will visit the calf several times per day to allow it to nurse (Galat-Luong 1981).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Galat-Luong 1981 source
Probably not a substantial claim: may not pass S3 (load-bearing). May combine two claims.
Premisenone givenVery little is known about C. liberiensis in the wild, either at the level of the individual or the population.
“Very little is known about Choeropsis liberiensis in the wild, either at the level of the individual or the population.”
Quote in the paperNot a substantial claimMay combine two claims.
PremisecitationThe species is highly dependent on streams and rivers, and is more widely distributed during the rainy season when water is more readily available.
“The species is highly dependent on streams and rivers, and is more widely distributed during the rainy season when water is more readily available (Robinson 1969(Robinson , 1970)).”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Robinson 1969
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Robinson 1970
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisereasoningCurrent assessments of wild C. liberiensis population numbers are largely unsubstantiated and based on qualitative data.
“Population numbers for wild Choeropsis liberiensis are unknown; current assessments are largely unsubstantiated and based on qualitative data.”
Quote in the paperNot a substantial claimProbably not a substantial claim: may not pass S3 (load-bearing).
Premisenone givenAll reports of disease in C. liberiensis stem from the captive population.
“All reports of disease in C. liberiensis stem from the captive population.”
Quote in the paperNot a substantial claimMay combine two claims.
PremisecitationC. liberiensis is primarily solitary; its favorite elements are forest and swamps, and its activity patterns are principally nocturnal with occasional activity at dawn and dusk.
“C. liberiensis is primarily solitary; its favorite elements are forest and swamps, and its activity patterns are principally nocturnal with occasional activity at dawn and dusk (Büttikofer 1890).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Büttikofer 1890
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
Probably not a substantial claim: may not pass S3 (load-bearing). May combine two claims.
PremisecitationAnother survey reported between 3.6 (primary rainforest) and 2.9 (secondary rainforest) C. liberiensis per km2, extrapolated to between 8,200 and 14,200 within Taï National Park and its buffer zones.
“Another survey based on radiotelemetry data for one female and extensive transect footprint counts reported between 3.6 (primary rainforest) and 2.9 (secondary rainforest) C. liberiensis per km 2 , and extrapolated to between 8,200 and 14,200 within Taï National Park and its buffer zones (Hentschel 1990).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Hentschel 1990
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationPopulation estimates for C. liberiensis range from 2,000 to 3,000 worldwide to possibly as many as 15,000 for Côte d'Ivoire alone.
“Population estimates range from 2,000 to 3,000 worldwide (Eltringham 1993(Eltringham , 1999;;Ransom et al. 2015) to possibly as many as 15,000 for Côte d'Ivoire alone (Roth et al. 2004).”
Quote in the paperNot a substantial claim0 of 3 sources checked- cites Eltringham 1993
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Ransom et al. 2015
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Roth et al. 2004 source
May combine two claims.
PremisecitationThe chorion of C. liberiensis is folded and its surface is covered with villi interspersed with rounded or irregularly shaped areolae; the villi are lined with columnar epithelium.
“The chorion is folded and its surface is covered with villi interspersed with rounded or irregularly shaped areolae; the villi are lined with columnar epithelium (Macdonald and Bosma 1985).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Macdonald and Bosma 1985 source
Probably not a substantial claim: may not pass S3 (load-bearing).
Premisenone givenIt is unknown if polycystic kidney disease is similarly prevalent in wild C. liberiensis populations.
“It is unknown if the condition is similarly prevalent in wild populations.”
Quote in the paperNot a substantial claimProbably not a substantial claim: may not pass S3 (load-bearing).
PremisecitationHybrids between C. liberiensis and the common hippopotamus do not occur.
“Hybrids between C. liberiensis and the common hippopotamus do not occur (Benirschke 2007).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Benirschke 2007
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationAnesthesia for C. liberiensis can be challenging; difficulties include accurate weight estimation, selection of protocols, tough skin, thick subcutaneous fat, limited access to peripheral circulation, endotracheal intubation, bradycardia, hypoventilation, apnea, and a high fatality rate.
“Anesthesia for C. liberiensis can be challenging; difficulties include accurate weight estimation for dosage calculations, selection of protocols for anesthetic drugs, tough skin that is challenging to penetrate with a needle or dart, a thick layer of subcutaneous fat leading to variable and systemic drug absorption, limited access to peripheral circulation, endotracheal intubation, bradycardia, h”
Quote in the paperNot a substantial claim0 of 2 sources checkedProbably not a substantial claim: may not pass S3 (load-bearing).
PremisecitationThe penis and prepuce of C. liberiensis curl caudally when not erect.
“The penis and prepuce curl caudally when not erect (Pocock 1923).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Pocock 1923 source
Probably not a substantial claim: may not pass S3 (load-bearing). May combine two claims.
InterpretationcitationThese assessments assume equal distribution of animals throughout all available areas of primary and secondary rainforest habitat and are likely to represent an overestimation of actual numbers.
“These assessments assume equal distribution of animals throughout all available areas of primary and secondary rainforest habitat and are likely to represent an overestimation of actual numbers (K.M. Hentschel, pers.comm.).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites K.M. Hentschel, pers.comm.
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationThe primary drivers of these land-use changes include increasing human population numbers, poor rural infrastructure and support services, and an often unstable political climate.
“The primary drivers of these land-use changes include increasing human population numbers, poor rural infrastructure and support services, and an often unstable political climate (Norris et al. 2010).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Norris et al. 2010 source
Probably not a substantial claim: may not pass S3 (load-bearing).
PremisecitationThe fetal placentas of three fullterm calves weighed 900-1,125 g.
“The fetal placentas of three fullterm calves weighed 900-1,125 g (Benirschke 2007).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Benirschke 2007
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationFlach et al. (1998) reported a heart rate of 42-100 beats/min and respiratory rate of 0-10 breaths/min for an anesthetized female C. liberiensis.
“Flach et al. (1998) reported a heart rate (beats/ min) of 42-100 and respiratory rate (breaths/min) of 0-10 for an anesthetized female.”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Flach et al. 1998 source
May combine two claims.
PremisecitationThere are two types of villi in C. liberiensis: areolar, which are more blunted, less branched, broader and overlie the endometrial glands, and interareolar villi.
“There are two types of villi: areolar, which are more blunted, less branched, broader and overlie the endometrial glands, and interareolar villi.”
Quote in the paperNot a substantial claimProbably not a substantial claim: may not pass S3 (load-bearing). May combine two claims.
PremisecitationThe diploid (2n) chromosome number for C. liberiensis is 36, with all chromosomes being metacentric except for an acrocentric Y chromosome.
“The diploid (2n) chromosome number for Choeropsis liberiensis is 36, with all chromosomes being metacentric except for an acrocentric Y chromosome (Hsu and Benirschke 1977).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Hsu and Benirschke 1977 source
May combine two claims.
PremisecitationOverall, the limited information in the primary literature describes C. liberiensis as a generally solitary species inhabiting primary and secondary rainforest, with preferred habitat types including low-lying Raphia palm swamps and areas in close proximity to or partially submerged in water.
“Overall, the limited information in the primary literature describes C. liberiensis as a generally solitary species inhabiting primary and secondary rainforest, with preferred habitat types including low-lying Raphia palm swamps and areas in close proximity to or partially submerged in water.”
Quote in the paperNot a substantial claimMay combine two claims.
PremisecitationMean values (n = 5 anesthetic episodes) from a juvenile female C. liberiensis were: pH 7.4; partial pressure of carbon dioxide 61.5 mmHg; partial pressure of oxygen 71 mmHg; bicarbonate 40.8 mmol/l; oxygen saturation 71.6%; end tidal carbon dioxide 42.5 mmHg; body temperature 36.3°C.
“Mean values (n = 5 anesthetic episodes) from a juvenile female were: pH 7.4; partial pressure of carbon dioxide (mmHg), 61.5; partial pressure of oxygen (mmHg), 71; bicarbonate (mmol/l), 40.8; oxygen saturation, 71.6%; end tidal carbon dioxide (mmHg), 42.5; and body temperature (°C), 36.3 (Morris et al. 2001).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Morris et al. 2001
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
Probably not a substantial claim: may not pass S3 (load-bearing).
InterpretationcitationThis activity is thought to comprise a form of territory marking or intraspecific communication.
“This activity is thought to comprise a form of territory marking or intraspecific communication (Lang 1975;Kranz 1982;Hentschel 1990;Robinson 1996;von Houwald et al. 2007).”
Quote in the paperNot a substantial claim0 of 3 sources checked- cites Lang 1975
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Kranz 1982 source
- cites Hentschel 1990
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationValues obtained from an anesthetized male C. liberiensis: average heart rate 60 beats/min; average respiratory rate 4 breaths/min; body temperature range 33.6-35.3°C; mean arterial hemoglobin saturation 95-99%.
“The following values were obtained from an anesthetized male C. liberiensis: average heart rate (beats/ min) 60; average respiratory rate (breaths/min) 4; body temperature range (°C) 33.6-35.3;mean arterial hemoglobin saturation 95-99% (Weston et al. 1996).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Weston et al. 1996
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationAllometric scaling was used to estimate the heart rate of conscious C. liberiensis at 60-65 beats/min; heart rate data for conscious animals has otherwise not been reported.
“Allometric scaling was used to estimate the heart rate (beats/min) of conscious animals at 60-65 (Bouts et al. 2012); heart rate data for conscious animals has otherwise not been reported.”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Bouts et al. 2012 source
May combine two claims.
PremisecitationA total of 120 animals, or two animals per linear kilometer of habitat, was reported from Azagny National Park based on transect footprint counts.
“A total of 120 animals, or two animals per linear kilometer of habitat, was reported from Azagny National Park based on transect footprint counts (Bülow 1987).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Bülow 1987
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationHabitat descriptions for 409 locations at Gola confirmed the presence of C. liberiensis in farm bush areas (54.41%), forest (24.21%), rivers/streams/swamps (16.14%), with one record from a cocoa plantation.
“Habitat descriptions for 409 locations based on search walks and camera traps at Gola Rainforest National Park, Sierra Leone, and Gola National Forest, Liberia, confirmed the presence of C. liberiensis in farm bush areas (243; 54.41%), forest (99; 24.21%), rivers, streams and swamps (66; 16.14%), with one record from a cocoa plantation (Hillers et al. 2016).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Hillers et al. 2016
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationThe multifocal areas of squamous metaplasia on both umbilical cord and amniotic membranes are unique to the hippopotamids and appear strikingly pustular.
“The multifocal areas of squamous metaplasia on both umbilical cord and amniotic membranes are unique to the hippopotamids and appear strikingly “pustular” (Macdonald and Bosma 1985;Benirschke 2007).”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Macdonald and Bosma 1985 source
- cites Benirschke 2007
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationThe size of wild C. liberiensis populations remains unknown but is widely considered to be declining based on the evidence of continued habitat loss and reduced densities reported for Côte d'Ivoire.
“The size of wild populations remains unknown but is widely considered to be declining based on the evidence of continued habitat loss and reduced densities reported for Côte d'Ivoire (Roth et al. 2004;Hoppe-Dominik et al. 2011).”
Quote in the paperNot a substantial claim0 of 2 sources checkedMay combine two claims.
PremisecitationC. liberiensis has diffuse epitheliochorial placentation and an umbilical cord with two arteries and two veins; however, a few specimens of C. liberiensis have only three umbilical vessels, two arteries and one vein.
“Choeropsis liberiensis has diffuse epitheliochorial placentation and an umbilical cord with two arteries and two veins (Macdonald and Bosma 1985); however, a few specimens of C. liberiensis have only three umbilical vessels, two arteries and one vein (Benirschke 2007).”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Macdonald and Bosma 1985 source
- cites Benirschke 2007
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationThe most commonly reported population estimate for C. liberiensis is ≤ 3,000; however, this figure originates from gray literature.
“The most commonly reported population estimate for C. liberiensis is ≤ 3,000; however, this figure originates from gray literature (Eltringham 1993(Eltringham , 1999;;Lewison and Oliver 2008).”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Eltringham 1993
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Lewison and Oliver 2008
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationEjaculates from seven C. liberiensis had an average volume of 3.0 ± 0.9 (SEM) ml and a concentration of 2.1 ± 1.3 × 10^6 spermatozoa per ml.
“Ejaculates from seven C. liberiensis had an average volume of 3.0 ± 0.9 (SEM) ml and a concentration of 2.1 ± 1.3 × 10 6 spermatozoa per ml (Saragusty et al. 2010).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Saragusty et al. 2010 source
May combine two claims.
PremisecitationThe only studies using radiotelemetry to evaluate home range size for C. liberiensis indicated an average female (n = 5) territory size of 40-60 ha and a male (n = 1) territory of 120 ha.
“The only studies using radiotelemetry to evaluate home range size for C. liberiensis indicated an average female (n = 5) territory size of 40-60 ha and a male (n = 1) territory of 120 ha (Bülow 1987;Hentschel 1990).”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Bülow 1987
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Hentschel 1990
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
Premisenone givenC. liberiensis forages primarily on dry land but does not rely on grass for its nutritional intake.
“Choeropsis liberiensis forages primarily on dry land but does not rely on grass for its nutritional intake.”
Quote in the paperNot a substantial claimMay combine two claims.
Premisenone givenMetabolite concentrations of progesterone in the saliva during the same time period were one-third to one-half of those in skin secretions, but the trends were comparable.
“Metabolite concentrations of progesterone in the saliva during the same time period were one-third to one-half of those in skin secretions, but the trends were comparable.”
Quote in the paperNot a substantial claimMay combine two claims.
PremisecitationVariables contributing to potential inaccuracy in population estimates include the inability to discern individual animals, the skill of detecting footprints and droppings, the decomposition rate of feces, and defecation rates.
“Variables contributing to potential inaccuracy include the inability to discern individual animals from camera trap footage or footprints, the skill of detecting footprints and droppings, the decomposition rate of the feces, and defecation rates (Bülow 1987;Roth et al. 2004).”
Quote in the paperSubstantial claim0 of 2 sources checked- cites Bülow 1987
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Roth et al. 2004 source
May combine two claims.
PremisecitationThe captive population at the end of 2015 (n = 377), believed to be derived from 60 founders, had an average inbreeding coefficient of 0.0600, an average mean kinship of 0.0305, and gene diversity of 0.9695.
“The captive population at the end of 2015 (n = 377), believed to be derived from 60 founders, had an average inbreeding coefficient of 0.0600, an average mean kinship of 0.0305, and gene diversity of 0.9695 (Steck 2016).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Steck 2016
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationThe reproductive anatomy of C. liberiensis has not been extensively detailed for either sex, although in many aspects it is likely to be similar to that of the common hippopotamus.
“The reproductive anatomy of C. liberiensis has not been extensively detailed for either sex, although in many aspects it is likely to be similar to that of the common hippopotamus (Macdonald 2007).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Macdonald 2007
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
PremisecitationData from 14 immobilized adult C. liberiensis were: mean heart rate 34 ± 7 beats/min; mean respiratory rate 14 ± 6 breaths/min; mean peripheral hemoglobin saturation 91-92% for animals receiving supplemental oxygen; mean end tidal carbon dioxide 55 ± 10 mmHg.
“Data from 14 immobilized adults were as follows: mean heart rate (beats/min ± SD) 34 ± 7; mean respiratory rate (breaths/min ± SD), 14 ± 6; mean peripheral hemoglobin saturation, 91-92% for animals receiving supplemental oxygen; mean end tidal carbon dioxide (mmHg ± SD), 55 ± 10 ( Bouts et al. 2012).”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Bouts et al. 2012 source
The quote only partly states this claim.
PremisecitationHigher birth weights are positively correlated with survival rate beyond 3 months of age.
“Higher birth weights were positively correlated with survival rate beyond 3 months of age.”
Quote in the paperNot a substantial claim0 of 1 sources checked- cites Leutenegger 1978 source
Probably not a substantial claim: may not pass S2 (checkable). May combine two claims.
InterpretationcitationC. liberiensis can serve as a flagship species for the Upper Guinean forest ecosystem, thus promoting sustainable and accountable land use; expansion, protection, and management of national parks and a highly viable platform on which to raise public awareness of this little known biodiversity hotspot.
“C. liberiensis can serve as a flagship species for the Upper Guinean forest ecosystem, thus promoting sustainable and accountable land use; expansion, protection, and management of national parks and a highly viable platform on which to raise public awareness of this little known biodiversity hotspot (Mallon et al. 2011;Fauna and Flora International and Forestry Development Authority 2013).”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Mallon et al. 2011
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Fauna and Flora International and Forestry Development Authority 2013
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
May combine two claims.
InterpretationcitationSex ratio varies among zoos, and thus husbandry factors associated with captivity are likely to play at least a partial role in the bias.
“Sex ratio varies among zoos, and thus husbandry factors associated with captivity are likely to play at least a partial role in the bias (Zschokke 2002;Boonyarittichaikij 2010;Flacke et al. 2015).”
Quote in the paperNot a substantial claim0 of 3 sources checkedMay combine two claims.
PremisecitationThe largest remaining legally protected, contiguous tract of Upper Guinean forest ecosystem lies within the Taï National Park and surrounding N'Zo Faunal Reserve; this area is believed to hold the largest remaining wild population of C. liberiensis.
“The largest remaining legally protected, contiguous tract of Upper Guinean forest ecosystem lies within the Taï National Park and surrounding N'Zo Faunal Reserve; this area is believed to hold the largest remaining wild population (Critical Ecosystem Partnership Fund 2015;Mallon et al. 2011).”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Critical Ecosystem Partnership Fund 2015
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Mallon et al. 2011
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
Probably not a substantial claim: may not pass S3 (load-bearing).
Premisenone givenIt is unknown if the female-biased sex ratio in captive C. liberiensis is an artifact of captivity or a reflection of normal biology for this species.
“It is unknown if the femalebiased sex ratio is an artifact of captivity or a reflection of normal biology for this species.”
Quote in the paperNot a substantial claimProbably not a substantial claim: may not pass S3 (load-bearing).
InterpretationcitationThe Krahn-Bassa/Sapo/Grebo/Taï complex spanning the border of Liberia and Côte d'Ivoire represents the greatest opportunity to establish and maintain protected areas with large intact stands of forest.
“The Krahn-Bassa/Sapo/Grebo/Taï complex spanning the border of Liberia and Côte d'Ivoire represents the greatest opportunity to establish and maintain protected areas with large intact stands of forest (Critical Ecosystem Partnership Fund 2015;Fauna and Flora International and Forestry Development Authority 2013).”
Quote in the paperNot a substantial claim0 of 2 sources checked- cites Critical Ecosystem Partnership Fund 2015
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
- cites Fauna and Flora International and Forestry Development Authority 2013
Couldn’t be matched to a paper in Crossref, so it can’t be checked.
The paper relies on this without support.
It is universally accepted that C. liberiensis numbers are limited.
“However, it is universally accepted that numbers are limited.”
The paper relies on this without support.
Zoos generally provide the entire ration for C. liberiensis as one or two daily feedings.
“in contrast, zoos generally provide the entire ration as one or two daily feedings.”
The paper relies on this without support.
Stress levels immediately prior to immobilization play an important role in the variability of success experienced with anesthesia protocols for C. liberiensis, especially when alpha-2 agonists are used.
“Stress levels immediately prior to immobilization play an important role in the variability of success experienced with these anesthesia protocols, especially when alpha-2 agonists are used.”
Self-check
Percentages, sums and reported differences re-computed from the paper’s own numbers.
Each contradiction between two passages listed here was confirmed by an independent check.
Nigerian taxon is called both a subspecies and a species
MinorThe paper recognizes heslopi as a subspecies of C. liberiensis, but later calls it the species C. heslopi without indicating a change in taxonomic treatment.
“Two subspecies of Choeropsis liberiensis are currently recognized (Lewison 2011): C. l. heslopi Corbet, 1969:388.”
“The Lower Guinean species, C. heslopi, was formerly reported in southeastern Nigeria, from the Owerri and Warri provinces of the Niger River Delta eastward from the Cross River (Ritchie 1930;Heslop 1945;Corbet 1969).”
Farm-bush percentage does not match the location count
MinorFarm-bush records account for 243 of 409 locations, or 59.41%, not 54.41%.
Reported 54.41 · recomputed 59.413203
“Habitat descriptions for 409 locations based on search walks and camera traps at Gola Rainforest National Park, Sierra Leone, and Gola National Forest, Liberia, confirmed the presence of C. liberiensis in farm bush areas (243; 54.41%), forest (99; 24.21%), rivers, streams and swamps (66; 16.14%), with one record from a cocoa plantation (Hillers et al. 2016).”
Eyeball measurements show greater width than height
MinorThe description says the eyeball is taller than it is wide, whereas its stated horizontal diameter, 24.3 mm, exceeds its vertical diameter, 22.5 mm.
“The globe (eyeball) itself is rounded and slightly taller than it is wide;”
“measurements from a subadult female (117 kg) were 24.3 mm horizontal diameter, 18.6 mm anterior to posterior diameter, and 22.5 mm vertical diameter (Hegner 1967).”
Wild male weight is reported, then excluded
MinorThe diagnosis gives a recorded weight for a wild adult male, but the general characters section says wild body-mass measurements exist only for two females.
“The only recorded weights for wild specimens are from an adult male at 204 kg (Heslop 1944) and two adult females at 165 and 170 kg (Hentschel 1990).”
“Morphometric measures, including body mass, are not available for any animals taken from the wild, except for the two adult females weighed by Hentschel (1990) during his fieldwork in Côte d'Ivoire.”
Cited juvenile blood-gas study concerns the other hippo species
MinorThe juvenile female's measurements are presented among C. liberiensis physiological values, but the cited reference identifies its subject as an East African river hippopotamus.
“Mean values (n = 5 anesthetic episodes) from a juvenile female were: pH 7.4; partial pressure of carbon dioxide (mmHg), 61.5; partial pressure of oxygen (mmHg), 71; bicarbonate (mmol/l), 40.8; oxygen saturation, 71.6%; end tidal carbon dioxide (mmHg), 42.5; and body temperature (°C), 36.3 (Morris et al. 2001).”
“114. Morris, P. J., et al. 2001. Chemical restraint of juvenile east African river hippopotamus (Hippopotamus amphibius kiboko) at the San Diego Zoo.”
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Cited by
Sorted by citations
Hart, L. J., Campione, N. E., & McCurry, M. R. (2022) · Palaeontology · doi:10.1111/pala.12629 · Cited by 9 · see citing works
“Likewise, E. megacephalus has a total length similar to the pygmy hippopotamus (Choeropsis liberiensis) and the above mass estimation also falls within a range akin to that of C. liberiensis (Flacke & Decher 2019).”
Avedik, A., & Clauß, M. (2023) · PLoS ONE · doi:10.1371/journal.pone.0291825 · Cited by 7 · see citing works
“Both species have a semi-aquatic lifestyle [15] and have evolved a complex forestomach system [16–18] where plant material is digested by the aid of a microbiome [19, 20].”
A new method to determine the diet of pygmy hippopotamus in Taï National Park, Côte d’Ivoire
Hendier, A., Chatelain, C., Du Pasquier, P., Paris, M., Ouattara, K., Koné, I., Croll, D., & Zuberbühler, K. (2021) · African Journal of Ecology · doi:10.1111/aje.12888 · Cited by 7 · see citing works
Predictors of animal sponsorship to support zoo‐based conservation activities
Doarks, T., & Arbuckle, K. (2024) · Conservation Science and Practice · doi:10.1111/csp2.13173 · Cited by 2 · see citing works
Kardos, M. C., Velmurugan, S., & Janečka, J. E. (2023) · Frontiers in Ecology and Evolution · doi:10.3389/fevo.2023.1144711 · Cited by 1 · see citing works
“, the pygmy hippopotamus averages 160–270 kg compared with the common hippopotamus averaging 1,400–1,500 kg) (Burnie and Wilson, 2005; Flacke and Decher, 2019).”
BackgroundInfluentialON SMALL CONTINENTAL MAMMOTHS AND DWARFISM
Kirillova, I., Markova, E. A., Панин, А. В., et al. (2023) · Зоологический журнал · doi:10.31857/s0044513423100045 · Cited by 1 · see citing works
Small Continental Mammoths and the Phenomenon of Dwarfism
Kirillova, I., Markova, E. A., Панин, А. В., et al. (2024) · Biology Bulletin · doi:10.1134/s1062359024700729
“A few cases of dwarfism have been described for wild animals, for example, the hippopotamus (Flacke and Decher, 2019) and the giraffe (Brown and Wells, 2020).”
Pérez‐Ramos, A., Moulaye, J., Martín‐Serra, A., et al. (2026) · Journal of Mammalian Evolution · doi:10.1007/s10914-026-09804-8
“In addition, a second specimen of Choeropsis liberiensis (BZC-CL-001) was obtained from the Brookfield Zoo Research and Vet - erinary Data Repository (Online Resource 1: Table S2).”
InfluentialGerakakis, N., Athanassiou, A., Kafousias, P., Anastasakis, G., & Lyras, G. (2026) · Journal of Mammalian Evolution · doi:10.1007/s10914-026-09824-4
- Authors
- Flacke, G. L., & Decher, J.
- Affiliations
- The University of Western Australia (AU)
- Zoological Research Museum Alexander Koenig (DE)
- Corresponding author
- Flacke, Gabriella L
- Venue
- Mammalian Species
- Publisher
- Oxford University Press (OUP)
- Published
- 2019-12-06
- Volume
- 51
- Issue
- 982
- Pages
- 100-118
- ISSN
- 0076-3519 (ISSN-L)
- 1545-1410
- OpenAlex
- W2992598458
- Semantic Scholar
- e006d656ab98e966a18d77a4b388c5e066759184
- Cited by
- 9
- References
- 101
- Citations by year
- 2026: 2 · 2024: 2 · 2023: 3 · 2022: 1 · 2021: 1
- FWCI
- 0.30
- Type
- journal article
- Language
- en
- Open access
- bronze
- doi.org
- Licence
- academic.oup.com/journals/pages/open_access/funder_policies/chorus/standard_publication_model (vor)
- Locations
- Mammalian Species (publishedVersion)
- Zenodo (CERN European Organization for Nuclear Research) (submittedVersion)
- Funders
- Rheinische Friedrich-Wilhelms-Universität Bonn
- Topics
- Physical Sciences › Environmental Science › Ecology › Wildlife Ecology and Conservation
- Health Sciences › Veterinary › Small Animals › Animal Behavior and Welfare Studies
- Social Sciences › Psychology › Social Psychology › Primate Behavior and Ecology
- Keywords
- Choeropsis liberiensis, pygmy hippopotamus, hippopotamus, threatened species, zoological collections, private collections, CITES Appendix II, conservation status
About these checks
Registry, retraction, Cites and statistics results are computed, not judged, with one exception: a printed reference that no index lists is read once by a model before it counts as a citation error. Claims are extracted by a model and kept only when their quote appears verbatim in the text; each cited source is then checked, with a model’s help, for whether it says what the claim relies on. Those verdicts can be wrong — read the claim’s quote and the cited source before relying on one.
Claims read by deepseek-v4.1-flash under definition version 3 (extract-v3) on 2026-10-27.
Classified by jev-1.13-20260917 (gold-guideline-v0.2) on 2026-10-27.
Cited sources are checked by gpt-6-sol.
Statistics and arithmetic are recomputed, not judged.
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